In 1963 Niko Tinbergen published a paper, “On Aims and Methods of Ethology,” dedicated to his friend Konrad Lorenz. This essay is a landmark in the development of ethology. Here Tm bergen defines ethology as “the biological study of behavior” and seeks to demonstrate the “close affinity between Ethology and the rest of Biology” (p. 411). Building on Huxley (1942), Tinbergen identifies four major problems of ethology: causation, survival value, evolution, and ontogeny. Cancern with these problems, under different names (mechanism, adaptation, phylogeny, and development), has dominated the study of animal behavior during the last half century (Dawkins, et al. 1991; Dewsbury 1992).