The engraved slate plaques were part of an extensive and variable class of ritual objects in Late Neolithic and Copper Age Iberia, with Classic plaques being the most numerous and standardized type. Classic plaques have a top and base separated by a horizontal line or bands, and base registers of repeating design elements (triangles, checkerboard, etc.). Associated with burials, they have been interpreted as genealogical records, with their base design referencing a clan or other social unit and their number of registers denoting the generational distance of the deceased from an important ancestor. The authors evaluate the genealogical hypothesis using a larger dataset than available when originally proposed, employing statistical analyses to examine the relationship between the number of registers and find locations, and between design elements and tomb size. Tomb size is viewed as a measure of collective labour, and hence a proxy of the status of the individuals in the tomb. These analyses show significant patterning between the number of registers and the plaques’ geographic distribution, and between specific design elements and tomb size, suggesting that the genealogical hypothesis remains a plausible explanation for the Classic plaques.

The dietary habits of Neanderthals are considered an issue of great interest in the literature and have opened an important number of fruitful debates. Indeed, understanding diets can provide important information regarding issues of palaeoenvironmental reconstructions and subsistence strategies. In this respect, dental remains can play a vital role in the conducted efforts to reconstruct the palaeoecological niches securely and accurately since dental microwear analyses have precisely detected dietary patterns of the populations in the past. In this context, the Iberian Peninsula forms an interesting model for examining Neanderthal populations, their subsistence strategies, and adaptive skills. This study aims the examination of already published data in order to provide a holistic approach regarding the dietary habits of H. neanderthalensis populations in the Iberian Peninsula, along with the importance of the utilization of dental microwear analysis in the archaeological record.

This study aims to determine the chronological sequence of the collective burials in the hypogea of the prehistoric cemetery of La Beleña (Cabra, Córdoba) through Bayesian analyses of 14C dates obtained from human remains. The data from this site are not only key to grasping the phenomenon of the introduction and spread of hypogea throughout the western Mediterranean, but to gain insight into multi-stage funerary practices during the Late Neolithic/Chalcolithic. The dataset comprises 14C dates of 71 of the 79 individuals placed in five of La Beleña’s six hypogea. The findings suggest: (i) La Beleña is one of the oldest assemblages of hypogea in Iberia, (ii) that this type of collective burial spread rapidly throughout the western Mediterranean area, (iii) that La Beleña is marked by two main phases of funerary activity interspersed by brief burial surges, (iv) funerary intensity at La Beleña increased between cal BC 3400–2900 (2σ), and (v) the cemetery saw a very brief surge of burials potentially related to a catastrophic event. The results of this analysis thus shed light on the little-known chronological sequence of prehistoric hypogea or rock-cut tombs in Iberia, their spread, and their relation to other Late Neolithic collective burials in western Europe.

Many European farmland bird populations are rapidly declining because of agricultural intensification and land-use changes. Robust estimates of population sizes and trends, habitat use, and protected area coverage within the distribution range are crucial to inform the conservation and management of threatened species. Here we report on the results of the 2019 Black-bellied Sandgrouse Pterocles orientalis (BBS) survey promoted and coordinated by SEO/BirdLife to update its breeding distribution, population size, and trends in continental Spain. A total of 660 grid cells, 10 × 10 km, Universal Transverse Mercator (UTM), were surveyed (81% of the distribution area), with 2,257 visits to 1,750 walked transects (7,001 km in total; 10.6 km per UTM). BBS was detected in 43% of sampled UTMs. At transect level, occupancy was 11% higher inside protected areas. At UTM level, occupancy was estimated at 0.58 (Bayesian credible interval [BCI] 95%: 0.55–0.61), revealing that BBS occupied about half of its previous breeding range (2003–2005). Using hierarchical distance sampling modelling, we estimated an average density of 1.33 individuals/km2 in occupied areas, and a population of 4,025 individuals (confidence interval: 1,840–7,609) within sampled areas, with an additional 697 individuals (confidence interval 461–1,075) in areas that were not surveyed. Further, the relative abundance of BBS (Kilometric Abundance Index) declined by 63% between 2005 and 2019 (annual decline rate of 4.5%). BBS used agricultural habitats (73%) and unprotected areas (54%) despite a higher occupancy within protected areas. Given the recent decline rate and persistent threats, the BBS conservation status should be upgraded to “Endangered” in peninsular Spain. Its future depends on land-use changes and agricultural practices, in particular the maintenance of fallows, semi-natural habitats, and pastures for extensive grazing. Better protection of important areas and targeted conservation initiatives should be promoted to halt and reverse the population decline in this key western Palearctic stronghold.

Research concerning transactions in the early first millennium bc in the westernmost Mediterranean has tended to focus on colonial coastlands occupied by scattered Levantine outposts, whereas cross-cultural interactions in hinterland regions have remained ill-defined. This article presents an assemblage of Egyptian vitreous artefacts, namely beads, a Hathor amulet, and further items from the seventh-century bc rural village of Cerro de San Vicente (Salamanca) in the interior of Spain. Macroscopic and chemical analyses demonstrate their likely manufacture in Egypt during the Middle and New Kingdom (second millennium bc), attesting to a far-reaching Phoenician maritime network that connected both ends of the Mediterranean. The authors interpret the items as liturgical objects, rather than mere high-status trinkets, that formed part of a widely shared Mediterranean world view and associated ritual mores. They consider the impact of cultural syncretism, which reached even remote and allegedly isolated peripheral settings in Iberia.

The spread of trapeze industries (the creation of trapeze-shaped flint tips) during Late Mesolithic is one of the most disruptive phenomena of technological change documented in the European Prehistory. Understanding the chronological patterns of this process requires (i) a critical evaluation of stratigraphic relationship between trapeze assemblages and radiocarbon samples, and (ii) considering different levels of chronological uncertainty according to the inbuilt age of the samples and the calibration process. In this paper, we critically evaluate and analyze the radiocarbon record of the first trapeze industries in the Iberian Peninsula. A dataset of 181 radiocarbon dates from 67 sites dated to 8800–8200 cal BP was collected and evaluated following a strict data quality control protocol, from which 135 dates of 53 sites were retained and classified according to a reliability index. Then, three different phase Bayesian chronological models were created to estimate the duration of the first spread of trapezes across Iberia, considering different levels of chrono-stratigraphic resolution. We find that trapeze industries appeared in the eastern half of Iberia, over an area of 330,000 km2 between 8505–8390 and 8425–8338 cal BP, spanning 0–85 yr (95.4% CI). When the oldest evidence of trapezes from Portugal are considered, the probability distribution expands (8943–8457 and 8686–7688 cal BP), due to the chronological uncertainty of human samples with marine diet and regional ΔR values applied. For the eastern half of Iberia, the current evidence indicates a very rapid spread of trapeze industries initiated in the Central-Western Pyrenees, suggesting cultural diffusion within Mesolithic social networks as the main driving mechanism.

Relatively few examples of Palaeohispanic writing have been recovered from the Vasconic territories of present-day Navarre, leading to the assumption that the Vascones were a pre-literate society. Here, the authors report on an inscription on a bronze hand recovered at the Iron Age site of Irulegi (Aranguren Valley, Navarre) in northern Spain. Its detailed linguistic analysis suggests that the script represents a graphic subsystem of Palaeohispanic that shares its roots with the modern Basque language and constitutes the first example of Vasconic epigraphy. The text inscribed on this artefact, which was found at the entrance of a domestic building, is interpreted as apotropaic, a token entreating good fortune.

Southern European peninsulas have repeatedly played notable roles as refuges in the natural history of flora during periods of glaciation. Euonymus latifolius (Celastraceae) is a relict species from the last Würmian glaciation in the Iberian Peninsula. It still lives with isolated populations in favourable, cool mountainous microhabitats, being an extremely rare, critically endangered species. These Iberian populations are often high-priority targets for conservation due to their long-term persistence and unique evolutionary trajectory. Previously, it has not been feasible to promote significant programmes for reintroduction and/or population reinforcement of this singular plant species due to the great difficulties of conventional propagation. In this study, we analysed the effects of temperature, light and gibberellic acid (GA3) on the germination responses of E. latifolius to develop an effective protocol for seed germination as a main outcome. The results are coherent with the climatic temperature conditions recorded broadly in the Iberian Peninsula in the past and in the current refuge locations for the taxon. The germination responses of E. latifolius are compatible with those of seeds with intermediate physiological dormancy. In particular, the seeds required a 10-week warm period (20/7°C + 15/4°C) followed by 20 weeks of cold period (5 + 1.5°C) to break dormancy and achieve germination values over 90%. GA3 also promoted germination (80%). Therefore, we developed the first effective protocol for promoting E. latifolius seed germination and, thus, sexual propagation, to facilitate urgent ex situ actions in the current climate change context.

La Mina is one of three sites, along with Cueva Millán and La Ermita, located in the middle course of the Arlanza river. La Mina was excavated for the first time in 2006 and three test pits were carried out. In one of them, evidence of two Palaeolithic occupations was identified and several remains of woolly rhinoceros were recovered. Amino acid racemisation dating yielded an age of 52.5 ka BP, the earliest Upper Pleistocene date for Coelodonta antiquitatis on the Iberian Peninsula. This new record may have several implications for understanding the access routes to the Castilian Plateau, together with the definition of a new migratory wave of this species at the end of the Pleistocene. The location of La Mina on the Castilian Plateau may help researchers to complete the movements of this species through the Middle and Upper Palaeolithic on the Iberian Peninsula.

Paradigms such as the coexistence of incineration and inhumation funerary practices in the northeast of the Iberian Peninsula during the Late Bronze Age are supported by the association of human remains with elements of material culture as guiding fossils. One example is the association established by Salvador Vilaseca in 1939 between the human remains and grooved pottery discovered in the Cova de Marcó in Tivissa (Ribera d’Ebre, Catalonia). This association has been accepted until today and even become a paradigm for the mixing of autochthonism (inhumation rites) with the introduction of material novelties such as grooved pottery and incineration rites during the first period of the Late Bronze Age. Direct radiocarbon (14C) dating of human remains from the Cova de Marcó shows that the remains originate from the Chalcolithic period. This indicates that there is no relationship between the sepulchral episode and the grooved pottery associated with it and used to date it. This disputes the paradigm regarding the coexistence of these two funerary practices during the Late Bronze Age and invites scholars to conduct a critical review using absolute dating techniques directly on the human remains of similar cases in order to verify or disprove the paradigm’s validity.

The well-documented versatility of donkeys (Equus africanus asinus) means that issues concerning their welfare can vary depending on how they are mainly used and their geographic location. The present study is the first assessment of donkey welfare to be systematically conducted in the Iberian Peninsula. This area is characterised by the coexistence of high levels of mechanisation and industrialisation, and human populations with low levels of formal education, still making use of donkeys in their daily work. This study aims to evaluate the main welfare problems affecting donkeys within this context. The welfare assessment was carried out in accordance with the first level of the AWIN protocol for donkeys. Only 37.6% of the animals evaluated showed all positive indicators, with no physical problem detected while the remaining 62.4% revealed at least one negative health indicator. Body Condition Score (BCS) was one of the main concerns, with around half of the donkeys exhibiting an inadequate body condition. Dental evaluation revealed similarly concerning results, with 62.8% of the animals assessed requiring treatment. Indications of hoof neglect were noted in 39.5% of the animals evaluated with a 9.84% incidence of lameness. Skin problems affected 26.7% of individuals. Only 41% of the animals displayed positive results for all behavioural indicators. Aside from BCS, the most prevalent problems were sub-optimal behaviour, pain reaction to cheek palpation, hoof disease and integument alterations. These issues should be prioritised, both as regards this specific donkey population and the education of their caretakers.

The Iron Age shipwrecks of Agde K (Brescou Island, France) and Cabrera B (Balearic Islands), discovered in the 1960s, together yielded seven lead ingots cast in the large shells of Pinna nobilis molluscs. Lead isotope analysis later traced the ingots to lead sources in south-eastern Iberia. These ingots are reassessed here as evidence for the integration of coastal production strategies in Iron Age south-eastern Iberia, revealing material connections between metallurgy and coastal industries linked to the exploitation of Pinna nobilis, such as sea silk manufacture. This compelling example of reuse of materials from one industry in another attests to a circular economic activity that is likely to have had practical and environmental motivations. The author aims to promote the recognition of Pinna nobilis shell casting and similar reuse phenomena elsewhere in the Mediterranean basin.

Large, conglomerate caves in north-eastern Iberia have been significant places since the Neolithic through to historical times; however, their significance during the Palaeolithic has barely been explored. This project is the first systematic study of the use of these iconic geological landmarks among Pleistocene hunter-gatherers.

The Iberian archaeological record is particularly rich in asynchronous (i.e. chronologically mixed) assemblages including Athenian pots that predate the other items by a couple of decades or even a few centuries. Recent scholarship on keimëlia, or ‘curated objects’ in modern parlance, has shown the potential of such objects to investigate questions of identity, agency and history-making among the receiving communities, but also to shed light on the role of Athenian pottery among them. This article analyses this phenomenon within the Iberian peninsula, focusing on drinking cups, both black-gloss with inset lip (Cástulo cups) and red-figure type B cups. Using case studies from necropoleis and settlements of the southeast and east of the peninsula, the article explores the reasons and meaning of this consumption practice. It is argued that the occurrence of ‘heirloom’ vases in Iberian tombs and their extraordinary survival in some settlements is the result of a conscious and deliberate choice indicating the existence of mechanisms of social distinction based on a diacritical use of material culture. It is further argued that different motivations might lie behind their delayed deposition: when the chronological gap between production and disposal dates is small, one or two human generations, curated Athenian vases worked similarly to non-curated ones, being emblematic of economic success, social affiliations and political rank. But when the interval is longer, Athenian pots became symbols of ancestry and elite status, possibly acquiring the same legitimizing role earlier bestowed upon Orientalizing artefacts. Supplementary material is available online (https://doi.org/10.1017/S0075426921000094) and comprises a catalogue of case-study objects.

The nematode genus Acrobeles is composed of two morphological groups distinguished by the presence (‘double’ cuticle) or absence (‘single’ cuticle) of the refringent inner layer of the cuticle. In the present study, four species of this genus, two with ‘single’ cuticle (Acrobeles ciliatus and Acrobeles cylindricus) and two with ‘double’ (Acrobeles aenigmaticus and Acrobeles complexus) are studied from coastal dunes in Spain. This study provides detailed morphological and morphometrical analyses for the four species, while molecular analysis, based on 18S and 28S ribosomal DNA, is provided for A. complexus. The four species are studied with scanning electron microscopy, which is obtained for the first time for A. cylindricus. These analyses revealed morphological and molecular differentiations between both groups, appearing as two related monophyletic entities. The subgenera Acrobeles and Seleborca, formerly considered as separate genera, are proposed to accommodate both groups.

Ectotherms are vulnerable to environmental changes and their parasites are biological health indicators. Thus, parasite load in ectotherms is expected to show a marked phenology. This study investigates temporal host–parasite dynamics in a lizard community in Eastern Spain during an entire annual activity period. The hosts investigated were Acanthodactylus erythrurus, Psammodromus algirus and Psammodromus edwardsianus, three lizard species coexisting in a mixed habitat of forests and dunes, providing a range of body sizes, ecological requirements and life history traits. Habitat and climate were considered as potential environmental predictors of parasite abundance, while size, body condition and sex as intrinsic predictors. Linear models based on robust estimates were fitted to analyse parasite abundance and prevalence. Ectoparasitic mites and blood parasites from two haemococcidian genera were found: Lankesterella spp. and Schellackia spp. Habitat type was the only predictor explaining the abundance of all parasites, being mostly higher in the forest than in the dunes. The results suggest that particularities in each host–parasite relationship should be accounted even when parasites infect close-related hosts under the same environmental pressures. They also support that lizard parasites can be biomarkers of environmental perturbation, but the relationships need to be carefully interpreted for each host–parasite assemblage.

Protected areas (PAs) are intended to preserve natural places, aiming to sustain ecosystem functionality and preserve biodiversity. However, PAs are spatially static, while major threats to biodiversity, such as climate and land-use change, are dynamic. The climatic conditions represented in a PA could vanish in the future and appear in other places more or less far away from the PA; these places could be considered as recipient areas potentially suited to receive propagules from the source PAs, which tend to lose the climatic conditions that motivated their protection. This study estimates the current and future climatic representativeness of mainland Iberian national parks by identifying future areas with a similar climate to those existing now in the parks and taking into account the degree of anthropogenic alteration and protection. We identify a network of ecological corridors connecting Iberian national parks with their recipient areas, as well as discriminating those most conflicting areas that impede network connectivity due to their degree of land-use transformation. Our results identify important areas for maintaining the climatic representativeness of Iberian national parks in the future, showing a substantial reduction in the climatic representativeness of the Iberian national parks. Although most of the recipient areas now have forest and semi-natural land uses and more than half of their whole area has protected status, current land uses in the Iberian Peninsula severely obstruct the corridor network connecting the parks and recipient areas.

Dryocosmus kuriphilus (Hymenoptera: Cynipidae) is a global invasive gall wasp and a pest of chestnuts (Castanea spp.). A study of the Chalcidoidea parasitoid community of D. kuriphilus was undertaken over two years, from March 2017 to March 2019, at 15 sites in south and northwest Spain (Málaga and Galicia regions). More than 18,000 galls were collected, and 1153 parasitoids belonging to 22 species of seven chalcidoid families, plus two individuals of an inquiline Cynipidae, Synergus facialis, emerged. Richness was higher in the Málaga region, with 20 species, while 17 parasitoids and one inquiline were identified in Galicia. The parasitism rate of native chalcid parasitoid species in both regions was low. Eupelmus urozonus and Mesopolobus lichtensteini were the most abundant native species. Mesopolobus tibialis was a dominant species in south Spain, while Ormyrus pomaceus was a dominant species in northwest Spain. Our results revealed the existence of a sub-community of univoltine, probably host specialized, parasitoids in south Spain, which overwinter in galls, exhibiting a similar life cycle to Torymus sinensis. These species were Torymus notatus, Aulogymnus bicolor, Aulogymnus obscuripes and Aulogymnus balani. Data on the recovery of T. sinensis after release in the south Spain region show it to be well established, but its numbers are still low in northwest Spain.