The genus Prolagus is very common in the Neogene and Quaternary of Europe, western Asia and northern Africa. However, the complexity of its systematics, mainly based on p3 characters that show great inter- and intraspecific variability, led to an under-utilization of Prolagus in biochronology and palaeogeography studies. A re-analysis of the species of Prolagus recorded in the Escobosa de Calatañazor karst fissure fillings (Duero Basin, northern Spain, MN7/8, late Middle Miocene) includes the introduction of new maxillary and mandibular morphological characters (position, shape and size of premolar and mandibular foramina) and new dental measurements (hypoflexus depth, distal hypercone length, and partial width of upper premolars; relative length of trigonid of lower molariform teeth) to be used for species discrimination within the genus Prolagus. The new morphological characters and measurements introduced in this paper allow for a more accurate taxonomic assignment of Prolagus species, thus improving the estimation of faunal palaeodiversity, detection of sympatric species and taphonomic mixing, and contributing to more reliable biochronological, palaeogeographical, and phylogenetic inferences based on Prolagus.

This paper presents an overview of the bryozoan fauna from the upper Koněprusy Limestone (kindlei Conodont Zone, middle Pragian, Lower Devonian) exposed in two quarries at Zlatý Kuň near Koněprusy in Central Bohemia, and discusses its paleoecology and paleobiogeography. The studied fauna is dominated by encrusting fistuliporine and trepostome bryozoans (eight species), accompanied mainly by reticulate fenestrates (four species), branching ramose trepostomes and cryptostomes (three species), and one massive trepostome species. The richest bryozoan association comes from reef core/margin facies (13 species), followed by crinoid-bryozoan facies of the ramp (eight species). The reef-terrace facies and the crinoid-bryozoan-algal facies contain three and two species respectively. Seven species are described taxonomically, three fistuliporines and four trepostomes. The following taxa are new: Koneprusiella armata n. gen. n. sp., Fistulipora rarivesiculata n. sp., Fistulipora hladili n. sp. and Leptotrypa varia n. sp. Paleobiogeographic patterns of the bryozoan fauna from the Koněprusy Limestone are similar to those of stromatoporoids, comprising widely distributed genera but mainly endemic species. This supports a relative geographic isolation of the Koněprusy reef. The bryozoan fauna from the Koněprusy Limestone shows paleogeographic affinities with that from the Lower Devonian (Pragian) of Morocco and the Middle Devonian of Michigan (USA).

Members of the subfamily Ursinae dispersed into North America from Africa and Asia during the Miocene, with the appearance of Ursavus (Schlosser, 1899), Indarctos (Pilgrim, 1913), and Agriotherium (Wagner, 1837) (Dalquest, 1986; Miller and Carranza-Castañeda, 1996; Hunt, 1998). However, none of these genera were thought to have survived past the Hemphillian Land Mammal Age in North America. It is thought that these genera were replaced, and possibly out-competed, by members of the extant genus Ursus (Linnaeus, 1758), or Plionarctos (Frick, 1926), as suggested by several sources (Bjork, 1970; Dalquest, 1986; Bell et al., 2004). It has also been suggested that the Ursavini (Agriotherium and Indarctos) may have given rise to the extant ursids and the Tremarctinae (Harrison, 1983; Miller and Carranza-Castañeda, 1996). Of the Ursavini, Agriotherium is consistently found in the Hemphillian Land Mammal Age, and so is used as an index fossil in that its absence is assumed to indicate that a site is Blancan rather than Hemphillian (Lundelius et al., 1987; Bell et al., 2004; Hunt, 2004).

Two taxa of the Late Carboniferous and four species of the Early Permian terrestrial snails have been found in the Late Paleozoic continental molasse sediments of the Upper Silesian-Cracow Upland (Southern Poland). Discovery of Anthracopupa ohioensis and Protodiscus priscus indicates that, besides in North America, they occurred also in the European part of the Pangea supercontinent. According to the general sedimentary facies context and the accompanying floral and faunal assemblages, the gastropods lived in swamp environments, including a topogenous fen.

The Actinoceramus sulcatus lineage (Parkinson, 1819) (Bivalvia: Inoceramidae) is a very distinctive and abundant component of late Albian (Early Cretaceous) molluscan assemblages that is found throughout Europe, Central Asia, Japan and the Far East of Russia, southern and western North America, South Africa, and possibly India, in a range of shallow- to deep-marine facies. The lineage encompasses a wide and continuous range of morphologies that provide evidence of phyletic evolution at varying rates combined with large ecophenotypic plasticity within populations. The evolution of A. sulcatus marks the oldest appearance of well-developed radial folds and sulci within the Inoceramidae. The range of morphological variation makes formal taxonomic subdivision of the group problematic. Here we use a combination of formal successional subspecies and informal morphotypes to subdivide the lineage into the following taxa: A. sulcatus forma sulcatus, A. sulcatus forma subsulcatus (Wiltshire, 1869), A. sulcatus forma munsoni (Cragin, 1894), and A. sulcatus biometricus Crampton, 1996. Within these taxa and morphotypes, we synonymise a large number of earlier names that have been applied to variants within the lineage. Each of the forms recognized has biostratigraphic utility and we describe four new lineage biozones, in ascending order: A. concentricus parabolicus, A. sulcatus, A. sulcatus forma munsoni, and A. sulcatus biometricus biozones. The lowest occurrence of A. sulcatus is approximately coincident with the base of the upper Albian as currently defined, at least throughout most of Europe, and this datum provides a valuable tool in correlation. The nature of radial folds within the A. sulcatus lineage poses interesting but still unanswered questions regarding shell morphogenesis in bivalves and the functional significance (if any) of radial folds in the Inoceramidae.

The furongian of Mount Kinnekulle, Västergötland, Sweden (Fig. 1) is exposed in a number of road cuts, stream cuts, and abandoned alum shale quarries scattered around the mountain (e.g., Westergård, 1922, fig. 18; Müller and Hinz, 1991, fig. 2). Stratigraphically, the exposed successions span the Olenus gibbosus Zone through the Peltura paradoxa Zone (Fig. 2), and they have a maximum thickness of ca 12 m (Westergård, 1943). The strata consist of interfingering layers of alum shale and bituminous limestone (colloquially referred to as “orsten”). Trilobites, predominantly olenids, occur frequently throughout the succession, although they are generally restricted to the limestones. During an investigation of trilobites in the Lotagnostus americanus Zone (Fig. 2) at two localities on Kinnekulle, the second author (JA) recovered two unusual trilobites. After a closer examination they could be assigned to Elkanaspis Ludvigsen, 1982 and Parabolinella Br⊘gger, 1882, both of which are extremely rare in the Furongian of Scandinavia.

Permian colonial corals from Artinskian to Kungurian strata in the Conglomerate Mesa area, Inyo Mountains, east-central California, include five new species, one of which is assigned to a new genus. The new taxa are: Malpaisia maceyi n. gen. and n. sp., Pararachnastraea bellula n. sp., P. delicata n. sp., P. owensensis n. sp., and Cordillerastraea inyoensis n. sp. These species, several of which compare most closely with other Artinskian and Kungurian species from eastern Nevada and northern Mexico, represent three distinct stocks that differentiated on an isolated submarine uplift offshore from the main part of the Cordilleran carbonate shelf.

Here we report the first detailed reconstruction of the eye and visual system of a Paleozoic ostracod, namely the dimorphic primitiopsid (Palaeocopa) Primitiopsis planifrons Jones, 1887. Evidence from the cuticular lens morphology and its position on the valve suggests that P. planifrons had a naupliar eye, which is the most common optical system in crustaceans. the characters of the eye, such as the separated cuticular lens on each valve and the strongly calcified nature of the lens and its diameter, are concordant with that of some Recent podocopid ostracods in which a divided type of naupliar eye is present. Ray tracing of the reconstructed cuticular lens of the primitiopsid species demonstrates strong spherical aberration and a very long focal length (as in the eye of podocopids), which implies the presence of a spherical mirror (tapetum) below the lens. Study of the ontogenetic change of the valve thickness aids the suggestion that P. planifrons was nektobenthic. the size of its cuticular lens in relation to that of living podocopid ostracods, and the water depth at which these Recent forms occur, indicates that P. planifrons may have lived in a relatively shallow, well lit environment such as the deep subtidal zone. These interpretations are consistent with supposed paleoenvironmental conditions derived from geological evidence. Thus, analysis of the nature and relative size of the eye gives an independent test to help identify paleoenvironmental parameters of long extinct ostracods.

Two brachiopod species of two of the earliest rhynchonelliformean families are described from one of the first studied Ordovician fossiliferous localities of northwestern Argentina. One is a new species of the family Euorthisinidae (Orthida, Plectorthoidea), Notorthisina musculosa Villas and Herrera. It displays peculiar features not observed in the family before, as well as a combination of internal and external features previously considered diagnostic of different genera, leading to emendment of the familial and generic diagnoses. Its dorsal muscle field, best impressed in large specimens, resembles that of Kvania Havlíček (Nanorthidae) in having four scars arranged radially. This feature reinforces the phylogenetic proximity of Kvania to the euorthisinids. The second studied species, Nanorthis bifurcata Harrington, has not been revised since its superficial description in 1937–1938, its type having been lost. Its description herein, based on a new collection from its type locality, allows a better knowledge of the group and helps clarify its supra-familial assignment.

Stromatomorpha californica Smith is a massive, calcified, tropical to subtropical organism of the Late Triassic that produced small biostromes and contributed in building some reefs. It comes from the displaced terranes of Cordilleran North America (Eastern Klamath terrane, Alexander terrane, and Wrangellia). This shallow-water organism formed small laminar masses and sometimes patch reefs. It was first referred to the order Spongiomorphidae but was considered to be a coral. Other affinities that have been proposed include hydrozoan, stomatoporoid, sclerosponge, and chambered sponge. Part of the problem was diagenesis that resulted in dissolution of the siliceous spicules and/or replaced them with calcite. Well-preserved dendroclone spicules found during study of newly discovered specimens necessitate an assignment of Stromatomorpha californica to the demosponge order Orchocladina Rauff. Restudy of examples from the Northern Calcareous Alps extends the distribution of this species to the Tethys, where it was an important secondary framework builder in Upper Triassic (Norian-Rhaetian) reef complexes. Revisions of Stromatomorpha californica produce much wider pantropical distribution, mirroring paleogeographic patterns revealed for other tropical Triassic taxa. Review of Liassic material from the Jurassic of Morocco, previously assigned to Stromatomorpha californica Smith var. columnaris Le Maitre, cannot be sustained. Species previously included in Stromatomorpha are: S. stylifera Frech (type species, Rhaetian), S. actinostromoides Boiko (Norian), S. californica Smith (Norian), S. concescui Balters (Ladinian-Carnian), S. pamirica Boiko (Norian), S. rhaetica Kühn (Rhaetian), S. stromatoporoides Frech, and S. tenuiramosa Boiko (Norian). Stromatomorpha rhaetica Kühn described from the Rhaetian of Vorarlberg, Austria shows no major difference from S. californica. An example described as S. oncescui Balters from the Ladinian-Carnian of the Rarau Mountains, Romania, is very similar to S. californica in exhibiting similar spicule types. However, because of the greater distance between individual pillars, horizontal layers, and the older age, S. oncescui is retained as a separate species. The net-like and regular skeleton of Spongiomorpha sanpozanensis Yabe and Sugiyama, from the Upper Triassic of Sambosan (Tosa, Japan), suggests a closer alliance with Stromatomorpha, and this taxon possibly could be the same as S. californica.

Deltaplax new genus, Deltaplax burdicki new species, and Deltaplax dellangeloi new species (Mollusca, Polyplacophora, Neoloricata, Multiplacophora) from the Mississippian Lower Buffalo Wallow Group (Chesterian) of Indiana, USA are described. the new genus is established by one partially articulated and one associated specimen with marginal fringes of two types of large spines, bilaterally symmetrical head and tail valves, and fifteen medial valves arranged in three longitudinal columns similar to those described previously for other multiplacophorans. the two specimens represent separate species differentiated by morphologies of the auxiliary valves, one type of spine, and subtrapezoidal versus triangular tail valves. the tail valve of the articulated specimen also had sutural laminae that projected under the preceding intermediate valve. the presence of sutural laminae allows for placement of multiplacophorans in Subclass Neoloricata of Class Polyplacophora. the head, intermediate, and tail valves are mucronate with comarginal growth lines and ridged insertion plates that probably inserted into soft tissue comparable to the girdle of modern polyplacophorans. the new specimens also indicate one left-handed, one auxiliary, and one right-handed valve in multiplacophorans was equivalent to a single bilaterally-symmetrical intermediate valve of extant polyplacophorans. However, multiplacophoran head valves have plates that project from the lower layer at the lateral margins and articulate with the first intermediate valves that overlap the head and second intermediate valves. These features have not been observed in more typical neoloricates, fossil or modern. Pending systematic revision of the class, Multiplacophora thus is retained as a separate order distinguished by the unique shared characters.

Within the Soom Shale Lagerstätte of South Africa (Upper Ordovician, Hirnantian), two brachiopod taxa preserve traces of organic tissue. In Trematis, presumed bands of periostracum are preserved on the flanks of the pedicle notch, and clay mineral casts of the pedicle are preserved in many specimens of Kosoidea. Both these genera are organophosphatic-shelled Linguliformea, together with a third genus identified as Plectoglossa. A fourth brachiopod taxon in the fauna belongs within the calcitic-shelled Rhynchonelliformea (Plectothyrella). Kosoidea cedarbergensis is a new species of discinoidean.

Abundant tubular macrofossils occur in finely laminated siltstones and shales of the 548–542 Ma Schwarzrand Subgroup, Nama Group, Namibia. The Nama tubes occur in both the Vingerbreek and Feldschuhhorn members commonly in dense populations and always in fine-grained, lower shore-face lithologies deposited below fair-weather wave base. The tubes are preserved mostly as compressed casts and molds that range in width from 0.6 to 2.1 mm; apparently incomplete specimens reach lengths up to 10 cm. All specimens show sinuous bending and occasional brittle fracture, indicating an original construction of strong but flexible organic matter. Feldschuhhorn specimens preserve fine longitudinal pleats or folds that record pliant organic walls, but the older Vingerbreek populations do not. Similarly, some specimens in the Feldschuhhorn Member display branching, while Vingerbreek tubes do not. The abundant Feldschuhhorn tubes are assigned to the widespread Ediacaran problematicum Vendotaenia antiqua; however, the distinctive Vingerbreek population remains in open nomenclature. The most abundant fossils in Nama rocks, these tubes resemble populations in Ediacaran successions from Russia, China, Spain, and elsewhere. Beyond their local importance, then, such tubes may turn out to be the most abundant record of Ediacaran life.

Two new well-preserved cornute stylophorans from the Upper Cambrian Whipple Cave Formation represent a new genus and species assigned here to Cardiocystella prolixora. These nearly complete specimens contain morphological information not available from other cornute specimens previously collected from this formation. Both specimens are preserved with superior faces exposed. One specimen contains a nearly complete theca but a somewhat disrupted aulacophore, whereas the other theca has been partially damaged by burrows but has a nearly complete but moderately eroded aulacophore. Cardiocystella prolixora exhibits wide marginals, abundant supracentral platelets, and an aulacophore with cover plates. Supracentral platelets cover much of the interior regions of the theca, which lacks a visible zygal bar in both specimens. In holotype 1791TX13, a bulge in the superior face of the theca likely shows the zygal bar position. The wide marginals of these new specimens resemble those of a specimen previously described from a partial theca and aulacophore assigned to Archaeocothurnus species indeterminate (Sumrall et al., 1997); however, the specimens described here are heart-shaped rather than boot-shaped. The placement of this specimen in a new genus and species is based on its unique marginal shape and arrangement.

X-ray CT scans at two different energies of three articulated specimens of Spathelopsis oakvalensis n. sp., a bivalve from the Bluefield Formation of the Mauch Chunk Group (Mississippian, Chesterian) from southeastern West Virginia, permitted observation and description of poorly known internal features of the shell, resulting in a reversal of the shell orientation alleged by past authors and concomitant reassignment of this problematic genus to the palaeotaxodont family Nuculanidae Adams and Adams. Distinctive posterior and anterior gapes in Spathelopsis suggest separation of inhalant and exhalant water streams, a characteristic associated with derived but not primitive palaeotaxodonts.

A new agelacrinitid edrioasteroid, Multiplexidiscus mckenziensis n. gen., n. sp. is described from the middle Silurian upper McKenzie Shale Member of the Mifflintown Formation of Pennsylvania. These edrioasteroids were attached to a large hardground clast that was reworked and flipped during a storm event. The occurrence of edrioasteroids in a near shore, perhaps in sub-normal salinity, is unusual for echinoderms. The edrioasteroids show a dispersed spatial distribution and weak thecal orientation. Multiplexidiscus differs from other agelacrinid edrioasteroids in the presence of a complex pattern of ambulacral cover plates combined with nearly straight paedomorphic ambulacra.

Ninety-five percent of the surface geology of The Netherlands consists of various Pleistocene sedimentary sequences. Of the other five percent, the principal area of ‘solid’ geology is in the south around Maastricht, in the province of Limburg, justifiably famous for its highly fossiliferous Upper Cretaceous succession, including the type section of the Maastrichtian Stage. Paleozoic exposures are very rare and, most relevant to the discussion herein, there is no exposed Permian succession. Yet the colonial history of The Netherlands makes it a haven for Permian researchers. The purpose of this brief communication is to alert interested researchers to the amalgamation of the Dutch Timor collections by the recent acquisition by the Nationaal Natuurhistorisch Museum - Naturalis, Leiden (NNM), of more than 10,000 specimens of Permian fossils, mainly marine invertebrates, from West Timor, Indonesia. Together with the collections already present at Naturalis, this easily forms the largest concentration of fossils from Timor in any museum.

Archamphiroa jurassica was described by Steinmann (1930) from carbonate deposits of the Cordillera de los Andes in Mendoza, central Argentina (Arroyo Negro, confluent of Malargüe River), assigned to the Callovian stage. Based on the general morphology and the internal structure of the identified fragments, Steinmann considered them to belong to the coralline algae. Comparisons made with some fossil and extant corallines species led him to the conclusion that Archamphiroa jurassica closely resembles some species of the extant genus Amphiroa.

Cambrian Cephalopods are presently reported only from tropical, carbonate platform successions that occur on a number of paleocontinents. Outside of West Gondwanan occurrences on the eastern Sino-Korean Platform in China, the record of Cambrian cephalopods is limited, and information on the early evolution and habitats of this molluscan class has grown slowly over the last century.

Three productidine brachiopods of Haydenella, Paraplicatifera, and Compressoproductus are recovered from the Tak Fa Formation (Wordian, Middle Permian) of the upper Saraburi Limestone Group exposed at Khao Wong of central Thailand (the western margin of the Indochina Terrane). The latter two genera are new to the Permian of Thailand, and the new species Paraplicatifera thaica is proposed herewith. Some taxonomic and nomenclatural problems in relation to the three genera are discussed. The assemblage suggests endemism for a Middle Permian marine faunule of the Indochina Terrane.