We study families of rational curves on irreducible holomorphic symplectic varieties. We give a necessary and sufficient condition for a sufficiently ample linear system on a holomorphic symplectic variety of $K3^{[n]}$-type to contain a uniruled divisor covered by rational curves of primitive class. In particular, for any fixed $n$, we show that there are only finitely many polarization types of holomorphic symplectic variety of $K3^{[n]}$-type that do not contain such a uniruled divisor. As an application, we provide a generalization of a result due to Beauville–Voisin on the Chow group of $0$-cycles on such varieties.

The base surface $B$ of a Lagrangian fibration of a projective, irreducible symplectic fourfold $X$ is shown to be isomorphic to $\mathbb{P}^{2}$.

We approach non-divisorial base loci of big and nef line bundles on irreducible symplectic varieties. While for K3 surfaces, only divisorial base loci can occur, nothing was known about the behaviour of non-divisorial base loci for more general irreducible symplectic varieties. We determine the base loci of all big and nef line bundles on the Hilbert scheme of two points on very general K3 surfaces of genus two and on their birational models. Remarkably, we find an ample line bundle with a non-trivial base locus in codimension two. We deduce that, generically in the moduli spaces of polarized K3[2]-type varieties, the polarization is base point free.