Ernietta plateauensis Pflug, 1966 is the type species of the Erniettomorpha, an extinct clade of Ediacaran life. It was likely a gregarious, partially infaunal organism. Despite its ecological and taxonomic significance, there has not been an in-depth systematic description in the literature since the original description fell out of use. A newly discovered field site on Farm Aar in southern Namibia has yielded dozens of specimens buried in original life position. Mudstone and sandstone features associated with the fossils indicate that organisms were buried while still exposed to the water column rather than deposited in a flow event. Ernietta plateauensis was a sac-shaped erniettomorph with a body wall constructed from a double layer of tubes. It possessed an equatorial seam lying perpendicular to the tubes. The body is asymmetrical on either side of this seam. The tubes change direction along the body length and appear to be constricted together in the dorsal part of the organism.

This study reports the earliest known reef-building anthaspidellid sponge, Rankenella zhangxianensis n. sp., from the Cambrian Series 3 (late Stage 5–early Guzhangian) deposit of the Zhangxia Formation, Shandong Province, China. Rankenella zhangxianensis mostly occurs within Epiphyton-Rankenella-Cambroctoconus reefs, with minor occurrence from inter-reef grainstone. The species has anthaspidellid-type regular ladderlike spicule networks consisting of dendroclones and trabs, and is characterized by trabs parallel/subparallel to the gastral surface that diverge and meet the dermal surface, which is typical of the genus. Compared to R. mors and R. hamdii, reported from the late Cambrian Series 2–late Cambrian Series 3 of Australia and the late Cambrian Series 3–early Furongian of Iran, respectively, R. zhangxianensis is characterized by a relatively thicker wall, high angle (~90°) between dermal surfaces and intersecting trabs, and minor occurrence of differentiated canals. On the other hand, R. zhangxianensis mainly shows obconical shape, which is far less diverse than the other two species showing conicocylindrical, digitate, explanate, or bowl shapes. These Cambrian Series 3 reefs from China are the ancestors of the Furongian anthaspidellid-microbial reefs and the Early Ordovician anthaspidellid-microbial reefs that flourished worldwide. They represent the resurgence of reef-building metazoans after the extinction of archaeocyaths at the end of Cambrian Series 2.

Transmitted light and scanning electron imaging of sectioned specimens of Conularia and Paraconularia, prepared using HCl etching and critical point drying, revealed that their periderm is composed of extremely thin (approximately 0.5–3 µm), variably distinct microlamellae that are alternately organic poor and organic rich. Organic-rich microlamellae are cross-connected by slender strands of organic matter originally embedded in calcium phosphate, which in etched specimens has been dissolved. Microlamellae may be organized in thicker (approximately 5–75 µm) layers, or macrolamellae, that vary in color and organic matter content, possibly owing to changes in the ambient paleoenvironment. Thickening of the periderm to form transverse ribs and internal carinae was achieved through gradual thickening of individual microlamellae. In the core of the transverse ribs and internal carinae the distinction between organic-rich and organic-poor microlamellae may be reduced, owing to organic material becoming dominant over (former) mineral matter or vice versa. Combined with observations of plicated aperture closure in thin-walled conulariids, including Archaeoconularia slateri (Reed, 1933) (Upper Ordovician, Scotland) showing smooth folding of midline carinae through angles greater than 90°, these results suggest a structure and original flexibility in the organic-rich biocomposite forming the conulariid periderm that supports its homology to the chitinous lamellar periderm of coronate scyphozoans.

A macrofloral assemblage dominated by elements of the Euramerican dryland biome is described from the Brazil Formation in Clay County, Indiana (Illinois Basin). Fossils were recovered from a thin heterolithic unit between a shallow-marine bed and the paleosol beneath the Minshall Coal, a Middle Pennsylvanian succession deposited near the Atokan-Desmoinesian and Bolsovian-Asturian boundaries. Sedimentological indicators imply accumulation under a seasonal climate, including interbedded siltstone and sandstone deposited during flashfloods, intraclasts eroded from local sources, and charcoal produced by wildfires. The macrofloral assemblage is consistent with a dryland setting, being dominated by large, coriaceous gymnosperm leaves with mesic to xeric traits, including Cordaites spp. indet., Lesleya sp. indet., and Taeniopteris sp. cf. T. multinervia. Sphenopsids and ferns typical of the wetland biome are rare. In contrast, the microfloral assemblage is dominated by fern spores, with lesser lycopsid spores and cordaitalean pollen. The succession indicates that the dryland biome predominated during late regression, prior to the onset of perhumid conditions that resulted in peat accumulation at late lowstand. However, the abundance of palynomorphs from wetland vegetation implies gradual fragmentation of the prevailing dryland flora and replacement by the wetland biome in the transition to glacial maximum. The taphonomic and paleobiogeographic context confirms that floras adapted to seasonal moisture deficit periodically dispersed into tropical lowlands, rather than being transported from ‘extrabasinal’ or ‘upland’ environments. The precocious occurrence of Taeniopteris, more typical of Late Pennsylvanian and Permian floras, may be the earliest record of the fossil-genus, and exemplifies the association of derived plant taxa with dryland habitats. The predominance of broad-leaved gymnosperms with mesic to xeric characters suggests that dryland communities contained more slow-growing and long-lived plants than contemporaneous wetland floras.

Small, stemmed frond fossils are common in Ediacaran-aged strata (ca. 565Ma) at Mistaken Point, Newfoundland, and many of them have previously been informally referred to as ‘dusters,’ but due to their small size and consequent relatively poor preservation, they have not yet been described taxonomically. Two new genera are herein defined on the basis of their unique constructions: the mop-like rangeomorph Plumeropriscum hofmanni new genus new species, and a flabellate, lobate frond Broccoliforma alta new genus new species. Plumeropriscum hofmanni n. gen., n. sp. has a three-dimensional petalodium structure with numerous primary branches attached at the base of the petalodium rather than at a central axis. Broccoliforma alta n. gen., n. sp. lacks visible branching, and instead has a lobate morphology with a petalodium that is at least superficially similar to the ivesheadiomorph Blackbrookia. Several other previously described taxa had also been included under the umbrella of ‘dusters.’ Collectively, these taxa show that the low epibenthic tier that small fronds occupied was more diverse than previously realized, with multiple taxa converging on the stemmed, small-frond body plan.

Taxonomic analysis is provided for a Middle Pennsylvanian macrofloral assemblage collected from clastic wetland deposits in Clay County, Indiana, on the eastern margin of the Illinois Basin. Adpressed plant fossils were recovered from four distinct beds in the lowermost Staunton Formation, positioned above the Minshall Coal (uppermost Brazil Formation), part of a succession deposited near the Atokan-Desmoinesian boundary. The assemblage of 22 fossil-taxa is dominated by pteridosperms (including Neuropteris flexuosa, Macroneuropteris scheuchzeri, Alethopteris densinervosa, Neuropteris ovata, Eusphenopteris neuropteroides, and Neuropteris missouriensis) with lesser cordaitaleans (Cordaites spp. indet.) and sphenopsids (particularly Sphenophyllum cuneifolium). Lycopsids are uncommon, and ferns are rare. In contrast, the microfloral assemblage from the Minshall Coal and overlying clastic units is dominated by lycopsid and tree fern spores. Comparisons with established biozonation schemes yield different ages depending on the regional biostratigraphic framework used: (1) latest Bolsovian (Radiizonates difformis Biozone, American microfloras); (2) latest Bolsovian or earliest Asturian (‘Neuropteris’ rarinervis Biozone, Appalachian Basin macrofloras); or (3) earliest Asturian (Linopteris obliqua Biozone, European macrofloras). The placement and correlation of the Bolsovian-Asturian and Atokan-Desmoinesian boundaries, which have traditionally been equated by palynology, are evaluated in the context of this discordance. Several revised stratigraphic scenarios are proposed for this interval in the Illinois Basin, which is being increasingly recognized as a time of significant environmental change throughout Euramerica. Homotaxial comparisons with European macrofloral assemblages indicate that, of the 18 biological taxa recorded, between 14 and 17 (78–94%) also are common in coeval wetland deposits in Europe. The similarities exemplify the spatial conservatism and low diversity of wetland plant communities over vast areas of tropical Euramerica, a manifestation of the intrinsically stressful conditions that characterize such habitats, and indicates that neither the Laurentian Shield nor the Appalachian-Variscan Mountains were an insurmountable barrier to plant dispersal during the Middle Pennsylvanian.

Three genera of gastropods described from the Lower Devonian Rockhouse Formation of Hardin County, Tennessee, currently are considered to be junior subjective synonyms of Platyceras (Visitator) Perner, 1911. Evaluation of the type specimens of these genera as well as types of two other gastropods from the same formation reveals the following: Platyostoma quadrangulare (Dunbar, 1920) is the only platyceratid gastropod among them; Saffordella tennessensis Dunbar, 1920 has a selenizone above midwhorl so it is an eotomarioidean belonging to the Gosseletinidae; Aulopea nelsoni Dunbar, 1920 and the lectotype of Distemnostoma princeps Dunbar, 1920 each has a broad subsutural sinus on the upper whorl surface similar to the Omphalotrochidae, as well as a basal sinus similar to certain primitive caenogastropods, so their systematic position is uncertain; I consider Holopea planidorsata Dunbar, 1920 to be a juvenile specimen of D. princeps; the fragmentary paralectotype of D. princeps possibly represents a gerontic form of Distemnostoma curtum Dunbar, 1920, which possesses a well-developed selenizone on an upwardly projecting shoulder at the edge of the upper whorl surface. Because D. curtum appears to be unrelated to D. princeps, the genotype of Distemnostoma, I propose the new generic name Omocordella for D. curtum. The new family Micromphalidae is erected to include Ordovician Slehoferia Rohr and Fryda, 2001, lower Carboniferous Micromphalus Knight, 1945, and Omocordella n. gen.

From new and more complete material, which includes frond fragments with casts of tracheid remains of the rachis, it can be demonstrated that the putative liverwort Hepaticites iporangae Ricardi-Branco, Faria, Jasper, and Guerra-Sommer, 2011, from the early Permian Rio Bonito Formation (Sakmarian) of the Paraná Basin, Brazil, is not a bryophyte but a tracheophyte. The new material was collected from the same locality and layer as the type material, in the Quitéria outcrop in the municipality of Encruzilhada do Sul, state of Rio Grande do Sul, Brazil. From the morphology of the deeply dissected pinnatifid pinnules with narrow laminar lobes, the taxon is provisionally reassigned to the genus Rhodeopteridium. Thus we propose the new combination ‘Rhodeopteridium’ iporangae new combination for this taxon. This new systematic interpretation contributes to our understanding of early liverworts (by removing Hepaticites iporangae as a possible taxon thereof) and clarifies an issue of diversity of the flora of the early Permian Rio Bonito Formation resulting from the original misidentification.

The ca. 780–740 Ma Chuar Group, Grand Canyon, Arizona, provides an exceptional record of life during the diversification of crown-group eukaryotes, just prior to the first Cryogenian glaciation. We document in detail the assemblage of organic-walled microfossils preserved in fine-grained siliciclastics throughout the unit. In contrast with earlier studies, we primarily used SEM to document fossil morphologies, augmented by transmitted light microscopy, FIB-SEM, and TEM. This resulted in the discovery of new species and the recognition of broad-ranging, intraspecific biological and taphonomic variation in other species. Twenty-two species and five unnamed morphotypes are described, including three new species: Kaibabia gemmulella, Microlepidopalla mira, and Volleyballia dehlerae; two new combinations: Galerosphaera walcottii and Lanulatisphaera laufeldii; and 17 previously described forms. The possible colonial green alga Palaeastrum dyptocranum Butterfield in Butterfield, Knoll, and Swett, 1994 and the index fossil Cerebrosphaera globosa (Ogurtsova and Sergeev, 1989) Sergeev and Schopf, 2010 (=C. buickii Butterfield, 1994) are described for the first time from Chuar rocks. Lanulatisphaera laufeldii, a locally abundant and globally widespread species characterized by submicrometer filamentous processes that form a reticulate network, may be a useful marker for the time interval just before the appearance of vase-shaped microfossils (VSMs) ca. 740 Ma.

Organic-walled microfossil assemblages decline in diversity upsection, coincident with the appearance of VSMs and intermittent euxinia within the basin. Whether this pattern is due to preservational bias related to greater water depth or the higher TOC of upper Chuar rocks or instead reflects biotic turnover related to the spread of euxinic water masses in the basin is unknown.

Cambrian biostratigraphy of the Indian subcontinent is best documented from the Parahio Formation of the Tethyan Himalaya. Recently established trilobite biostratigraphy shows that the formation encompasses the latest part of unnamed Stage 4 and much of unnamed Stage 5. A variety of small shelly fossils have been recovered via acid digestion of carbonate beds and include tetract and pentact hexactinellid sponge spicules, chancelloriid spicules belonging to Chancelloria sp. and a new species, Archiasterella dhiraji, shells of an helcionelloid comparable to Igorella maidipingensis, a meraspid ptychopariid trilobite, the tubular Cupitheca sp., a poorly preserved hyolith, and an assortment of spinose microfossils of uncertain affinity. These newly recovered microfossils are consistent with the trilobite-based lower and middle Cambrian age determination and do not support a late Cambrian age for the top of the Parahio Formation advocated in some recent literature. The microfossils reported herein significantly expand the known diversity of such fossils from Cambrian strata in the Himalayan region, and allow for comparison of this fauna with others from Gondwanaland and elsewhere. Integration with trilobite data indicate that the stratigraphic ranges of many small shelly fossils described in this study are greater than previously recognized.

Clusters of associated colony fragments discovered weathering out of bedding planes in the Upper Ordovician of the Cincinnati, Ohio, region provide a rare opportunity to quantify intracolony variation in ramose stenolaemate bryozoans. Sixteen colonies were reassembled as completely as possible from 198 fragments, and the following colony-level characters were measured: colony dimensions, branch link length and diameter, and branch order. Results indicate that branch link length and diameter systematically decrease as colonies grow via branch bifurcation. Branching ratio (i.e., the number of distal first-order branches divided by the number of immediately proximal second-order branches) appears to be more genetically than environmentally controlled and to be consistent among orders of stenolaemates and perhaps across the phylum. Colonies with endozones mined out by endoskeletozoans result in broken branches as opposed to pristine growing tips. This varies stratigraphically, perhaps in response to the distribution of the boring animals. The rarity of borers and the systematic proximal increase in branch diameter in these colonies suggest the zooids in the proximal portions of the colonies were alive at the time of colony death. If the time and effort can be invested in reassembling colonies, these morphometric data can then be applied to taxonomic, phylogenetic, and paleoenvironmental studies.

Catenipora is one of the most common tabulate coral genera occurring in various lithofacies in the Upper Ordovician Xiazhen Formation at Zhuzhai in South China. A combination of traditional multivariate analysis and geometric morphometrics is applied to a large number of specimens to distinguish and identify species. Based on three major principal components extracted from 11 morphological characters, three major groups as determined by the cluster-analysis dendrogram are considered to be morphospecies. Their validity and distinctiveness are confirmed by discriminant analysis, descriptive statistics, and bivariate plots. Tabularium area and common wall thickness are the most meaningful characters to distinguish the three morphospecies. Geometric morphometrics is adopted to compare the morphospecies with types and/or figured specimens of species previously reported from the vicinity of Zhuzhai. Despite discrepancies in corallite size, principal component analysis and discriminant analysis, as well as consideration of overall morphological characteristics, indicate that the morphospecies represent C. zhejiangensis Yu in Yu et al., 1963, C. shiyangensis Lin and Chow, 1977, and C. dianbiancunensis Lin and Chow, 1977.

Catenipora occurs in seven stratigraphic intervals in the Xiazhen Formation at Zhuzhai, representing a variety of heterogeneous environments. The coralla preservation is variable due to differential compaction; coralla preserved in limestones are commonly intact and in growth position, whereas those in shales are mostly crushed or fragmentary. The size and shape of corallites are considered primarily to be species-specific characters, but are also related to the depositional environments. In all species, morphological characters, including corallite size, septal development, and shape and size of lacunae, show high variability in accordance with lithofacies and stratigraphic position. The intraspecific differences in corallite size at various localities in the Zhuzhai area may indicate responses to local environmental factors, but may also reflect genetic differences if there was limited connection among populations.

The Campanian rudist fauna identified from the localities Playa Panamá, Santa Rosa National Park, Colorado de Abangares, and Bolsón in Guanacaste Province, Costa Rica, is composed of the antillocaprinids Antillocaprina sp. aff. A. suboccidentalis Chubb, 1967, Antillocaprina sp. and Antillocaprinidae indet.; the multiple-fold hippuritids Barrettia monilifera Woodward, 1862, Parastroma trechmanni Chubb, 1967, and cf. Whitfieldiella gigas (Chubb, 1955); the plagioptychids Plagioptychus trechmanni Chubb, 1956, Plagioptychus sp. cf. P. zansi Chubb, 1956, Mitrocaprina costaricaensis new species, Mitrocaprina multicanaliculata Chubb, 1956, and Plagioptychidae indet.; and the radiolitids Radiolites sp. aff. R. multicostata (Adkins, 1930), Alencasterites new genus mooretownensis (Trechmann, 1924), Bournonia? tetrahedron (Chubb, 1967), Chiapasella sp., Guanacastea costaricaensis new genus new species, cf. Potosites sp. aff. P. tristantorresi Alencáster and Pons in Pons et al., 2010, and aff. Thyrastylon sp. Although some differences in rudist diversity and sedimentary setting were observed among localities, most species indicate the same age for all occurrences, which correspond to the Campanian, probably mid-Campanian. Radiolitid specimens appear better preserved than those of other taxa and are thus discussed more in detail. Both the number of radiolitid genera exclusively known from the New World, and reasonable doubts about the correct attribution of some New World species to Old World genera, indicate important differences between rudist faunas at both sides of the Atlantic (Mediterranean and American Tethys, respectively) during the Late Cretaceous.

Angiosperms first appeared in the fossil record as pollen during the Valanginian–Hauterivian; they spread out of the tropics in the Aptian and Albian, and radiated in the Late Cretaceous. Despite these general patterns, details of the taxonomic, geographic, and ecological evolution of Cretaceous angiosperms are relatively poorly known because only a handful of Early and mid-Cretaceous macrofloras have been reported. This is the first detailed report of a fossil leaf flora from the Cedar Mountain Formation from the mid-Cretaceous of the Western Interior. We describe a flora that is overwhelmingly dominated by angiosperms (152 of 153 identified specimens are angiosperms) from the Albian–Cenomanian transition that is preserved in a clay- and carbonate-rich, lacustrine mudstone from the uppermost Cedar Mountain Formation of Emery County, Utah. We recognize 18 leaf morphotypes, all of which are dicotyledonous angiosperms. The majority of the Cedar Mountain morphotypes have taxonomic affinities with forms of similar age described from the Atlantic and Gulf coastal plains and other localities from the Western Interior. From this, we infer that a relatively diverse angiosperm flora grew along the margins of a small pond on the coastal plain. Palynological preparations of the fossil matrix were barren; however, previous studies of other facies within the formation showed that both conifers and ferns were important components of the regional vegetation during Cedar Mountain time. The effective absence of conifers and ferns in this macroflora and low leaf mass per area values among the angiosperms measured suggests that even at the Early–Late Cretaceous transition, angiosperms had come to dominate some sites, particularly those that were disturbed or seasonally ephemeral, where fast-growth or seasonal deciduousness would have been favored.

The isolation of Zealandia in the latest Cretaceous and early Paleogene following the final break-up of Gondwana fostered significant provincialism in molluscan faunas, concomitant with the segmentation of oceanic circulation patterns and changing climate nearing the end of the greenhouse phase in the Southwest Pacific. The earliest volutes (Gastropoda: Volutidae) from the Zealandian region reflect this isolation after separation from the Marie Byrd Land region of West Antarctica with several endemic groups being recognized for the first time since collecting first started in the Wangaloa Formation at Mitchells Rocks (Wangaloa), South Island, New Zealand in 1869. Five taxa attributed to the Volutidae are described herein from South Island (Wangaloa Formation and Steel Greensand) and also the Chatham Islands (Red Bluff Tuff) from the early (mid-Danian) to late Paleocene (Thanetian?). These comprise a new mid-Danian fulgorariine? genus and species, Wangaluta henaconstricta n. gen. n. sp.; a new combination also from the Wangaloa Formation, Wangaluta? neozelanica (Finlay and Marwick, 1937); two new zidonine volutes, the mid-Danian Alcithoe. s.l. wangaloaensis n. sp. from the Wangaloa Formation, and also Teremelon onoua n. sp. from the late Paleocene-early Eocene of the Red Bluff Tuff; and a probable volute, Fulgorariine? gen. indet. sp. indet. from the mid-Danian of the Steel Greensand. All of these taxa are endemic in the early Paleogene of New Zealand and represent a significant boost in our knowledge of post-K-Pg boundary diversification of volutid gastropods in the shrinking Weddellian Biotic Province in the southern rim of the Pacific.

A new monospecific “caninioid” genus, Preisingerella n. gen., from the lower Permian of the Karavanke Mountains (Southern Alps, Slovenia) is erected. The type species is Preisingerella stegovnikensis n. sp. The new taxon can be differentiated from other Caninia-type genera by its specific ontogeny and features of its dissepimentarium. Corals with such morphology had a wide distribution during the Carboniferous and early Permian, occurring in mostly shallow-water carbonate rocks. The phylogenetic relationships within this group are mostly unclear due to similarities in the adult stages. The earlier stages reveal the main distinguishing features that are decisive for a generic assignment, but these have rarely been well preserved and properly considered. The new taxon is compared with related genera of the Cyathopsidae and species of Caninella Gorskiy, 1938 characterized by lateral dissepiments. Large numbers of specimens of the new species, representing a monospecific assemblage, have been collected from the Born Formation at Mt. Stegovnik. Sedimentological and microfacies characteristics, as well as macro- and microfossil assemblages, underline this correlation. The fusulinoidean assemblage of the Born Formation, with Sphaeroschwagerina carniolica (Kahler and Kahler, 1937), as the predominant species, corresponds to the time span between the Sphaeroschwagerina moelleri-Schwagerina fecunda and Pseudofusulina moelleri zones, indicating a late Asselian to early Sakmarian age in the Southern Urals.

Estimates of Precambrian eukaryotic diversity and disparity indicate broad trends of increase in the Mesoproterozoic Era, leading to a peak and then rapid decline by ca. 750 Ma. The organic-walled microfossil assemblage presented here is representative of that mid-Neoproterozoic height of eukaryotic species richness. Organic-rich shales and siltstones of the mid-Neoproterozoic upper Alinya Formation, eastern Officer Basin, Australia, preserve an abundant and diverse assemblage of organic-walled microfossils deposited in a low-latitude, shallow marine setting. Use of scanning electron microscopy (SEM) revealed an unexpected level of morphological detail not visible in transmitted light microscopy. This led to the recognition of new species as well as establishment of degradational sequences, which aid in fossil recognition. In total, 26 taxa are described here; these include 21 previously named forms, four newly described species (Caelatimurus foveolatus, Culcitulisphaera revelata, Karenagare alinyaensis, and Morgensternia officerensis), and one new combination (Vidalopalla verrucata).

Zigzagopora wigleyensis n. gen. n. sp. is an Upper Ordovician (Sandbian, early Caradoc) cyclostome bryozoan from the Arbuckle Mountains of Oklahoma, USA. It has runner-type colonies characterized by a mostly uniserial, geniculate arrangement of monomorphic zooids that bud alternately left and right, producing a zig-zag pattern of growth. This new genus has calcified interior walls and non-pseudoporous exterior walls. It is thus most likely affiliated with the paleotubuliporine family Sagenellidae, despite superficial similarities with the corynotrypid cyclostomes with which it co-occurs.

The recently described radiodontan Lyrarapax unguispinus Cong et al., 2014 from the Chengjiang biota (Cambrian Series 2, Stage 3) highlighted a new morphological type of frontal appendage and unique mouth structures, a functional combination reinforcing the diversification of feeding strategies of radiodontans during the early Cambrian. Here we describe Lyrarapax trilobus n. sp. from the same fossil Konservat-Lagerstätte. The new species differs from L. unguispinus in the morphology and distribution of endites on the frontal appendage and the strengthening structure of the body flaps. The two species resemble each other in body shape (pattern of flap size), neck segment number, cephalic plates, and most importantly a mouth characterized by concentric wrinkled furrows. The latter confirms that a soft mouth without sclerotized plates is a real feature of Lyrarapax and supports the idea that oral structures provide valid diagnostic characters within Radiodonta.

The early Albian to mid-Albian cosmopolitan genus Douvilleiceras is represented in the Haida Gwaii archipelago of western British Columbia by five species, D. mammillatum (Schlotheim), D. offarcinatum (White), D. scabrosum Casey, D. spiniferum (Whiteaves), and D. aff. spiniferum (Whiteaves). Specimens of one of these taxa, D. spiniferum, are particularly abundant on Haida Gwaii and the species is discussed with respect to its morphological variability and ontogenetic development. The important morphological features for taxonomic criteria of the genus Douvilleiceras are the mode of tuberculation, the pattern of ribbing, and the proportions of the shell in the middle growth stage.