A latest Paleocene charophyte flora collected from the South Gobi area in the Junggar Basin, western China, includes the geographically widespread taxa Peckichara torulosa var. varians (Dollfus and Fritel, 1919) Sanjuan, Vicente, and Eaton, 2020, Lychnothmanus vectensis (Groves, 1926) Soulié-Märsche, 1989, and Gyrogona lemani capitata Grambast and Grambast-Fessard, 1981. Lychnothmanus vectensis (as Lychnothmanus aff. L. vectensis) is known from the Cretaceous–Paleocene transition in eastern China and the latest Paleocene in western China, with likely additional records from the United States (Utah). The earliest European records of L. vectensis are from the late Eocene to early Oligocene in Spain, France, and England. Similarly, the oldest record of G. lemani capitata is from the latest Paleocene in the South Gobi area, with younger records from the middle Eocene of France. These latest Paleocene gyrogonite assemblages demonstrate the origin of these charophyte lineages in Asia. The dispersal of these charophytes from Asia to Europe in the middle to late Eocene appears to have occurred before the retreat of the Turgai Strait in both the Tarim area and the Siberian Basin by the end of the late Eocene and before the “Grande Coupure” in Europe and the Mongolian Remodelling in Asia during the Eocene–Oligocene transition. We hypothesize that waterbirds may have facilitated this intercontinental dispersal, and that idea is supported by the shared occurrence of avian groups in Central Asia and Europe in the middle and late Eocene.

Modular coral-like fossils occur in thrombolitic reefal beds at two stratigraphic levels within the Lower Ordovician (Floian) Barbace Cove Member of the Boat Harbour Formation, in the St. George Group of western Newfoundland. They are here assigned to Reptamsassia n. gen.; R. divergens n. gen. n. sp. is present at both levels, whereas a comparatively small-module species, R. minuta n. gen. n. sp., is confined to the upper level. Reptamsassia n. gen. resembles the Ordovician genus Amsassia in its phacelocerioid structure, back-to-back walls of adjoining modules, module increase by longitudinal fission involving infoldings of the wall, tabula-like structures that are continuous with the vertical module wall, and calices with concave-up bottoms. The new genus is differentiated by its encrusting habit, modules with highly variable growth directions and shapes throughout skeletal growth, and modules that may separate slightly or diverge from one another following fission. Together, Amsassia and Reptamsassia n. gen. are considered to represent a distinct group of calcareous algae, the Amsassiaceae n. fam., which possibly belongs to the green algae. The Early Ordovician origination of Amsassia followed by Reptamsassia n. gen. contributed to the beginning of the rise in diversity on a global scale and in reefal settings during the Great Ordovician Biodiversification Event. Reptamsassia minuta n. gen. n. sp. was an obligate symbiont that colonized living areas on its host, R. divergens n. gen. n. sp., with isolated modules of R. divergens n. gen. n. sp. able to persist in the resulting intergrowth with R. minuta n. gen. n. sp. This is the earliest known symbiotic intergrowth of macroscopic modular species, exemplifying the development of ecologic specialization and ecosystem complexity in Early Ordovician reefs.

A new species of the Paleozoic bryozoan genus Ptilotrypa of the order Cryptostomata is described from the lower part of the Yong Limestone Formation, Katian, Upper Ordovician of the Kumaun Tethys Himalaya: Ptilotrypa bajpaii new species. The presence of the genus Ptilotrypa in the Tethyan Himalaya suggests paleogeographic connections to the Upper Ordovician of North America and, consequently, Upper Ordovician age for the lower part of the Yong Limestone Formation. This species displays a reticulate colony shape, which suggests an efficient filtering capacity in an environment with a high primary production. Morphological peculiarities and systematic assignment of the genus Ptilotrypa are discussed.

UUID: http://zoobank.org/898276c8-2924-4da2-ae96-3392cb2ebbc3

The Lochkovian, Pragian, and basal part of the Emsian, which represent the post-Kwangsian Orogeny strata in the South China Block, are mainly composed of siliciclastic rocks. This lithological composition impedes investigation of Pragian and Lochkovian conodont biostratigraphy in the South China Block, which results in a persistent controversy on the age of relevant lithological units. The present study provides new evidence by reporting for the first time Lochkovian conodonts obtained from the South China Block, specifically the Gaoling Member of the Nahkaoling Formation at the Lingli section, central Guangxi. The conodont fauna, consisting of Pandorinellina exigua lingliensis Lu n. subsp., Pandorinellina exigua exigua, Zieglerodina? tuojiangensis Lu n. sp., Amydrotaxis praejohnsoni, and Eognathodus cf. E. irregularis, places the studied interval of the Gaoling Member in the lower or middle Lochkovian (contingent upon varying definitions for the base of the middle Lochkovian) to lower Pragian. Moreover, Amydrotaxis praejohnsoni, which was reported previously only in North America and eastern Australia, is herein also recorded in the South China Block, and thus may play an important role in intercontinental biostratigraphical correlation. By shedding light on the age of the upper limit of the underlying Lianhuashan Formation at the Lingli section, the present study indicates that the Kwangsian Orogeny ended before the late Lochkovian. This date is slightly earlier than the previously estimated late Lochkovian based on studies of fossil plants from the siliciclastic rocks deposited after the Kwangsian Orogeny.

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The Cretaceous chaetetid sponge Acanthochaetetes huauclillensis Sánchez-Beristain and García-Barrera is reported for the first time from the Aptian–Cenomanian Khalsi Formation, Ladakh Himalaya, India. Its low- to high-domical growth form could suggest an adaptation to either an environment with constant sedimentation rates, or to an irregular substrate. However, these growth forms also may indicate an absence of important environmental/sedimentological changes during the lifespan of the sponges. In addition, the growth form of this species suggests a calm, non-turbulent, reef-like microenvironment. Along with the other faunal assemblages, such as the rudists, corals, and the gastropod Nerinea, A. huauclillensis indicates a tropical to subtropical shallow marine carbonate platform setting. This new finding extends its stratigraphic range from the upper Hauterivian to the Aptian–Cenomanian interval in the eastern Tethyan realm.

Borophagus is the terminal genus of the highly diverse and successful subfamily, Borophaginae. Skeletal remains of this bone-crushing canid are most commonly found in transitional or grassland environments across North America between Late Miocene–Middle Pleistocene, but are rare or absent in forested habitats. Here, we describe a humerus from the Gray Fossil Site of Tennessee, which is the first occurrence of this genus in a heavily forested ecosystem. The distinct limb proportions of Borophagus suggest the genus may have been well suited for a closed habitat like the Gray Fossil Site, contrary to where a majority of their fossils have been previously found. This discovery documents the first pre-Pleistocene occurrence of a canid in the Appalachian region of the eastern United States.

Closterocrinus elongatus Hall, 1852, long assigned to the cladid inadunate crinoid family Ampheristocrinidae (=Euspirocrinidae), has been found to possess pinnulate biserial arms and a cup composed of four exsert basals, five radials, a primanal above the radial circlet, and fixed primibrachials and secundibrachials. Accordingly, this species is herein reassigned to the monobathrid camerate crinoid family Paramelocrinidae.

Three new species of the shallow-marine, warm-water bivalve Plicatula are reported from the upper Paleocene Santa Susana Formation of southern California. Plicatula simiensis new species is from the middle part of the formation on the south side of Simi Valley and occurs as a displaced specimen in deep-marine turbidites. Plicatula lapidicina new species and P. trailerensis new species are both from coralline-algal-rich muddy siltstone just beneath a nearshore, coralline-algal limestone interval in the upper part of the formation in the Santa Ynez Canyon area, east-central Santa Monica Mountains. These three new species represent the first late Paleocene records of genus Plicatula on the west coast of North America and the first Paleocene records of this genus in southern California.

West Falls Group strata (Chemung facies) of Upper Devonian (Frasnian) age in New York State reflect sedimentation on a storm dominated sandy shelf. Rugose corals occur in storm deposited coquinites rich in bioclastic debris and are abundant locally, although representing only three species, Tabulophyllum orientale (Stumm), Macgeea ponderosa Stumm, and Disphyllum caespitosum (Goldfuss). The Tabulophyllum and Macgeea species are both characterized by large amounts of stereome (biogenic calcite) in the apical part of the corallite, perhaps useful in stabilizing corallites in the sediment.

Nacocrinus elliotti new genus, new species, is reported from the beta member of the Naco Formation (Pennsylvanian, Desmoinesian) at the Kohl Ranch locality in central Arizona. This is the first pachylocrinid reported from the Naco Formation.

The silurian graptoloid genus Globosograptus was erected by Boucek and Přibyl (in Přibyl 1948, p. 37), who selected as type species Monograptus wimani Bouček, 1932. As noted by Bulman and Rickards in the Treatise (Bulman 1970, p. V155), Globosograptus “was based on the silhouette appearance of a slender Llandovery species on which the thecae possess a long prothecal portion, no thecal overlap, and a seemingly enrolled methathecal portion. The nature of this enrollment is indefinite … The genus has no precise significance and its application is quite impracticable.”

The editor of the Zoological Record has kindly informed me that the generic name Polysiphonia Finks, 1995, (class Calcarea, order Murrayonida, family Maeandrostiidae) (Finks, 1995, page 4) is preoccupied (Mrs. M. Joan Thorne, personal commun., 1996). I therefore rename it Polysiphonaria.

Triangulotarbus terrehautensis n. gen. and sp. is a Pennsylvanian phalangiotarbid (Arachnida) that occurs in the sandy shales of the lower Shelburn Formation, McLeansboro Group, Desmoinesian Series, above Coal VII (Danville Coal) of the Dugger Formation. Triangulotarbus terrehautensis is the youngest phalangiotarbid that has been collected in North America.

A specimen of the unstalked Cretaceous crinoid Uintacrinus socialis Grinnell, 1876, has a theca with an anomalous appearance that may be attributed to either inaccurate regeneration or unusual taphonomy. The former would entail regeneration of brachials from pinnulars, and inaccurate regeneration of the arms themselves. The latter would imply that the theca was broken into at least three pieces, and then the pieces were rotated with respect to each other. The presence of broken plates and sediment in the line of breakage indicates that the specimen was broken after death, and the taphonomic explanation is favored. Of the hundreds of known specimens of this species, this is the only one with such an appearance.

Twenty-four species of pteropods are described from the Pliocene Miyazaki Group in the Tsuma-Takanabe district, Miyazaki Prefecture, southwest Japan. Five are new: Clio shibatai new species, Cuvierina miyazakiensis new species, Cavolinia floridana japonica new subspecies, C. itoigawai new species, and C. vendryesiana hyugaensis new subspecies.

The pteropod fauna of the two lower members of the Miyazaki Group consists largely of extinct species. Whereas more than one-half of the species of the two upper members are extant.

Twelve species, Limacina sp., Clio hataii (Noda, 1972), C. shibatai new species, Cuvierina miyazakiensis new species, Bowdenatheca? sp., Diacria digitata subsp. indet., Cavolinia floridana japonica new subspecies, C. cf. C. gypsorum (Bellardi, 1873), C. itoigawai new species, C. mexicana (Collins, 1934), C. rattonei Simonelli, 1896, and C. vendryesiana hyugaensis new subspecies seem to serve as good index fossils for the Pliocene.

The extensive fauna of vertebrates from the Lower Devonian Water Canyon Formation of Utah was initially reported and described by Branson and Mehl (1930, 1931), and subsequently described in detail by Denison (1952, 1953, 1958, 1967). A major part of the fauna consists of heterostracans, particularly pteraspidids (Denison, 1953, 1967), and to date six species are known from the Water Canyon Formation. This fauna comes from the upper Emsian and is similar in age to that of the Beartooth Butte Formation of Wyoming and Montana, and also to faunas recently reported from Nevada and California (Elliott, 1989; Ilyes and Elliott, 1991, 1992).