Two new species of petrified seeds Eosperma edromense, and Anasperma burnense gen. et sp. nov. are described from the Cementstone Group of Berwickshire. Anasperma shows that the anatropous condition existed among Palæozoic ovules; it has a single integument with two lateral apical lobes.

Rachides of Stauropteris berwickensis sp. nov. and associated megasporangia are also described.

An account is given of the theory of the telomic origin of the first ovular integument. The second (outer) integument in Angiosperm ovules may therefore have evolved after the establishment of anatropy, either from the first integument or as an overgrowth of the chalaza. Evidence is cited to support the theory that Angiosperm carpels have evolved from dorsiventral bivalved cupules.

Among most of the geologists who had of late years been engaged in investigating the structure of the North-West Highlands, and especially among those who did not concur in Murchison's explanation of the phenomena exhibited there, it was a growing belief that great overthrusts had been largely instrumental in producing the remarkable stratigraphical relations of the rock masses of that region. After a most careful detailed examination of the ground by the Geological Survey, the existence and importance of such thrusts was not only placed beyond a doubt, but a variety of additional remarkable structures were discovered, which open up new fields of investigation to the physical geologist.

Glaucophane-Schists and eclogite are associated with ultrabasic rocks, basic intrusions and Ordovician spilitic lavas at Knockormal in the Girvan-Ballantrae complex, South Ayrshire. Detailed mineralogical and chemical data are presented which indicate that the various glaucophane and blue-amphibole rocks have been derived from spilitic lavas. In addition to glaucophane a blue-amphibole related to crossite is also described. While there is no evidence for an introduction of soda, the deformation attending the glaucophanitic metamorphism was accompanied by chemical changes which appear to have occurred at high water-vapour pressures. There is no evidence of any relationship between the eclogite and adjacent glaucophane-rocks. The eclogite, which is highly aluminous, is interbanded with wehrlite, pyroxenite and serpentinite, and represents a particular phase in the crystallization of these ultrabasic rocks.

The rise of archosaurs during the Triassic and Early Jurassic has been treated as a classic example of an evolutionary radiation in the fossil record. This paper reviews published studies and provides new data on archosaur lineage origination, diversity and lineage evolution, morphological disparity, rates of morphological character change, and faunal abundance during the Triassic–Early Jurassic. The fundamental archosaur lineages originated early in the Triassic, in concert with the highest rates of character change. Disparity and diversity peaked later, during the Norian, but the most significant increase in disparity occurred before maximum diversity. Archosaurs were rare components of Early–Middle Triassic faunas, but were more abundant in the Late Triassic and pre-eminent globally by the Early Jurassic. The archosaur radiation was a drawn-out event and major components such as diversity and abundance were discordant from each other. Crurotarsans (crocodile-line archosaurs) were more disparate, diverse, and abundant than avemetatarsalians (bird-line archosaurs, including dinosaurs) during the Late Triassic, but these roles were reversed in the Early Jurassic. There is no strong evidence that dinosaurs outcompeted or gradually eclipsed crurotarsans during the Late Triassic. Instead, crurotarsan diversity decreased precipitously by the end-Triassic extinction, which helped usher in the age of dinosaurian dominance.

A detailed account of the morphology and ontogeny of the late Middle Cambrian crustacean †Henningsmoenicaris scutula is presented. Ten successive ontogenetic stages could be recognised in the material collected from various localities in Sweden. Morphogenetic changes include the development of a pair of stalked lateral eyes and the increase in the number and size of appendages and their setal armature. Notably, early stages lack ‘proximal endites’ on all post-antennular appendages; such a spine-bearing endite has previously been thought to appear simultaneously on these limbs. In †H. scutula a single functional endite appears on the third limb in an advanced stage; an additional endite appears on the second limb and, subsequently, further endites appear on more posterior limbs. Furthermore, a single specimen of †Sandtorpia vestrogothiensis gen. et sp. nov. is described. Based on this new information and data of other ‘Orsten’ taxa, particularly those assigned already to the early evolutionary lineage of Crustacea, a small-scale computer-based phylogenetic analysis was performed. This resolved the basal branchings of Crustacea s. l. as follows: †Oelandocarididae (=†Oelandocaris oelandica+†H. scutula+†S. vestrogothiensis)+(†Cambropachycopidae (=†Goticaris longispinosa+†Cambropachycope clarksoni)+ (†Martinssonia elongata+Labrophora (=†Phosphatocopina+Eucrustacea))). Plotting ontogenetic data on the phylogram and comparing the ground pattern at every node led to the detection of three peramorphic heterochronic events in the evolutionary lineage towards Eucrustacea.

In Part I brief descriptions are given of the goniatite faunas found at horizons in the Scottish Carboniferous succession; and mainly on the evidence of these faunas, the stages of B, P1; P2, E1; and E2 and A are approximately demarcated. No goniatites suggesting the presence of H, R and G have been seen.

In Part II descriptions are given of certain species of the following genera: Beyrichoceratoides, Anthracoceras, Cravenoceras, Cravenoceratoides, Girtyoceras, Eumorphoceras, Goniatites, Sudeticeras, Dimorphoceras, Neodimorphoceras and a new genus, Cluthoceras.

There is abundant evidence of injury, teratology and pathological conditions in trilobites although it is commonly difficult to distinguish between their morphological results. Many injuries were probably sustained during moulting, with spines, bilamellar fringes and narrow gaps between dorsal exoskeleton and doublure being most vulnerable. Some injuries were the result of predaceous attack and provide important clues to the trophic structure of Palaeozoic communities. Many injuries show evidence of repair. Some supposedly teratological conditions, most notably the number of paradoxidid thoracic segments, are so common that they must be considered part of normal variation. Others represent genetic mutation or ontogenetic malfunction, the latter possibly externally induced. Undoubted pathological conditions are restricted to swellings and borings presumably caused by parasitic infestation.

Abnormalities affecting lateral areas are more common than those involving the rachis and glabella and this probably reflects both greater mortality when the axial region is involved, and greater vulnerability of peripheral regions to predaceous and moulting damage. The first explanation also accounts for the rarity of cephalic abnormalities compared to those of the pygidium.

High Precision U-Pb zircon and monazite dating in the Aiguilles Rouges–Mont Blanc area allowed discrimination of three short-lived bimodal magmatic pulses: the early 332 Ma Mg–K Pormenaz monzonite and associated 331 Ma peraluminous Montées Pélissier monzogranite; the 307 Ma cordierite-bearing peraluminous Vallorcine and Fully intrusions; and the 303 Fe-K Mont Blanc syenogranite. All intruded syntectonically along major-scale transcurrent faults at a time when the substratum was experiencing tectonic exhumation, active erosion recorded in detrital basins and isothermal decompression melting dated at 327-320 Ma. Mantle activity and magma mixing are evidenced in all plutons by coeval mafic enclaves, stocks and synplutonic dykes. Both crustal and mantle sources evolve through time, pointing to an increasingly warm continental crust and juvenile asthenospheric mantle sources. This overall tectono-magmatic evolution is interpreted in a scenario of post-collisional restoration to normal size of a thickened continental lithosphere. The latter re-equilibrates through delamination and/or erosion of its mantle root and tectonic exhumation/erosion in an overall extensional regime. Extension is related to either gravitational collapse or back-arc extension of a distant subduction zone.

In 1804, Lieutenant-Colonel Imrie communicated to the Royal Society of Edinburgh a paper entitled “A Description of the Strata which occur in ascending order from the Plains of Kincardineshire to the Summit of Mount Battoc, one of the most elevated points in the Eastern District of the Grampian Mountains.” He pointed out that, in a short stretch of six miles in the North Esk section, “we pass from the secondary horizontal strata of the newest formation to the vertical, contorted primary strata of the oldest date, and terminate with granite, the primitive rock in the conception of many geologists.”

The Schichallion complex is situated in the Central Highlands of Scotland between the villages Struan and Kinloch Rannoch. The area is of considerable geological interest and has been investigated stratigraphically and structurally by E. M. Anderson and Bailey and McCallien. As a consequence of their researches a complete stratigraphical succession has been established. Thus, the metamorphic rocks of the area are classified into the Moinian and Dalradian systems, which are separated by a plane of tectonic discontinuity known as the Boundary Slide. The Moinian rocks are quartz-felspathic granulites, whereas the Dalradian system includes pelitic schists, quartzites, limestones as well as a variety of meta-igneous rocks.

In his previous research the present author has established the tectonic history of the complex. In particular three episodes of folding (F1 to F3) and a much later episode of faulting (F4) were recognized. Of these the F3 episode is of least significance. Consequently, events after the F2 movements can be in many cases regarded as post-folding.

The present contribution is concerned essentially with the mapping of the metamorphic zones (garnet and staurolite-kyanite) and with a detailed study of the mineralogical evolution of regionally metamorphosed rocks within these zones.

In the field it is possible to prove that the staurolite-kyanite zone is essentially post-folding, since the kyanitepegmatites are found to cut across the minor F2-folds. In this respect textural studies confirm the field observations. The study of the internal inclusions in garnets indicates that the garnet zone has a much longer history, since pre-F2 garnets are found in the southern part of the complex and throughout the central part of the area syn-tectonic F2 garnets are apparent. The pre-F2 garnets contain a very fine-grain F1 fabric. Hence the garnets in relation to the F1 movements are post-tectonic. Thus, the regional metamorphism can be subdivided into three phases: the F1 metamorphism, the F2 metamorphism and the post-F2 metamorphism. The latter, on structural evidence appears to be at least in part contemporaneous with the F3 movements.

The meta-igneous rocks of the area are grouped into the hornblende-schists and granular epidiorites. The hornblende-schists appear to have suffered deformation and recrystallization during F1 and F2 episodes of movement. On the other hand the granular epidiorites are later than the F1 and the F2 movements. Although in the southern parts of the district the epidiorites have been slightly deformed, elsewhere they preserve the original ophitic texture and cut across the F1 and F2 folds. The deformation in the south is attributed to the effects of the F3 folding. The epidiorites have been evidently emplaced as dolerites after the F2 movements and before the F3 metamorphism.

The localized retrogressive metamorphism is associated with the F4 movements, which are responsible for the Loch Tay Fault. The Fault is later than the minor intrusives associated with the Younger Granites of the Scottish Highlands and is suggested to be of a Lower or Middle O.R.S. age.

On the basis of the chemical composition of the plagioclase felspars it is proposed to include all the staurolite and kyanite bearing rocks into the epidote-amphibolite facies. In this respect temperature and the hydrostatic pressure are assumed to have been the main factors, since similar minerals came into existence during static and dynamic stages of metamorphism alike.

The state of Texas has one of the greatest records of pterosaurs in the world, surpassing all other US states and most countries in the number of occurrences. Uniquely, this record extends over the entire 150+ million history of the Pterosauria. A review of this pterosaur record confirms at least 30 pterosaurs known from 13 occurrences, including five valid species. The holotypes of two of these species have been described before and are diagnosed and erected here as the new species Radiodactylus langstoni, gen. et sp. nov., named in honour of Dr. Wann Langston Jr, the father of Texas pterosaurology, and Alamodactylus byrdi, gen. et sp. nov.. Phylogenetic analysis of all Texas pterosaurs that can be coded for more than one character confirms that these species are distinct from others and occupy phylogenetic positions close to their original classifications. Radiodactylus langstoni is recovered as a non-azhdarchid azhdarchoid, Quetzalcoatlus northropi as an azhdarchid, Alamodactylus byrdi as a non-pteranodontoid pteranodontian, Aetodactylus as a pteranodontoid, and Coloborhynchus wadleighi as an ornithocheirid. The presence of eudimorphodontid, dsungaripterid, as well as other azhdarchid and pteranodontoid pterosaurs, is also confirmed in Texas.

Several mesosocopic structures are consistent with mechanical accumulation of crystals and movement of melt in granite magmas, as well as compaction and shear of crystal-melt aggregates, concentrations of microgranitoid enclaves indented by megacrysts, and concentrations of crystals of the same mineral with different crystallisation histories. Evidence for crystal and enclave accumulation is shown clearly in mafic and silicic layered intrusions (MASLI-type granite plutons), for example, the Kameruka Granodiorite, Bega Batholith, south-eastern Australia.

Crystal accumulations with interstitial liquid may become mobile in a magma chamber, owing to instabilities in the host magma caused by seismic and replenishment events or thermal and buoyancy variations. This remobilised material may intrude other parts of the chamber, as well as earlier-formed cumulates and even wall-rocks, as dykes, tubes, troughs and pipes. Marked concentrations of accessory and mafic minerals may also develop in these structures. Interstitial melt may also be extracted from accumulated aggregates, intruding and disrupting the aggregates. Spectacular examples of these various structures are preserved in the Tuolumne Batholith, California. Detailed mechanisms for the formation of many of the structures are not well understood, though the formation of cumulates in vertical layers suggests that sorting and filter pressing during flow and resulting strain of crystal mushes may play important roles.

Eusuchians with deep snouts and labiolingually compressed teeth are known from the Palaeogene of Laurasia. These are usually referred to Pristichampsinae, but the type species, Pristichampsus rollinati, is based on insufficiently diagnostic material and should be treated as a nomen dubium. At least two Lutetian species formerly referred to Pristichampsus can be recognised – Boverisuchus magnifrons in Germany and possibly elsewhere in Europe, and Boverisuchus vorax, new combination, in western North America. Material from the middle Eocene of Italy and Texas may represent distinct species. A phylogenetic analysis confirms their close relationship and also supports a relationship with two Asian forms – early Eocene Planocrania datangensis and Palaeocene Planocrania hengdongensis. The name Planocraniidae Li 1976 is applied to this group. A distinctive quadrate with a prominent dorsal peak between medial and lateral hemicondyles is known only in Boverisuchus, and although the teeth of Planocrania are flattened, they are not serrated. Planocraniids maintain a phylogenetic position as the sister group to Crocodyloidea+Alligatoroidea, but this part of the tree is unstable and discovery of older, more primitive planocraniids will help resolve conflicts on the phylogenetic relationships of extant crocodylian lineages.

The Watch Hill Grits, in the lower Ordovician Skiddaw Group of the Lake District, are dated as latest Tremadoc or earliest Arenig (older than the Didymograptus deflexus Biozone) by means of acritarchs and graptolites. This places them lower in the Skiddaw sequence than previous authors supposed, and they are thought to have been thrust southwards over younger Arenig rocks. The interval inferred for the Watch Hill Grits brackets the Tetragraptus approximatus Biozone and its equivalents in Australasia, S China, N America and Scandinavia. The dating of the Watch Hill Grits suggests that equivalents of the Tetragraptus approximatus Biozone are present in N England and that sedimentation continued in the Lake District during a period of global eustatic regression. Other beds in the Lake District and on the Isle of Man have yielded similar acritarch assemblages and are probably correlatives. Five new acritarch taxa are described: Caldariola gen. nov., Acanthodiacrodium? dilatum, Stellechinatum sicaforme and Striatotheca prolixa spp. nov., and Tetraniveum arenigum cumbriense subsp. nov.; one new combination, Caldariola glabra (Martin) comb, nov., is proposed.

Several mesosocopic structures are consistent with mechanical accumulation of crystals and movement of melt in granite magmas, as well as compaction and shear of crystal-melt aggregates, concentrations of microgranitoid enclaves indented by megacrysts, and concentrations of crystals of the same mineral with different crystallisation histories. Evidence for crystal and enclave accumulation is shown clearly in mafic and silicic layered intrusions (MASLI-type granite plutons), for example, the Kameruka Granodiorite, Bega Batholith, south-eastern Australia.

Crystal accumulations with interstitial liquid may become mobile in a magma chamber, owing to instabilities in the host magma caused by seismic and replenishment events or thermal and buoyancy variations. This remobilised material may intrude other parts of the chamber, as well as earlier-formed cumulates and even wall-rocks, as dykes, tubes, troughs and pipes. Marked concentrations of accessory and mafic minerals may also develop in these structures. Interstitial melt may also be extracted from accumulated aggregates, intruding and disrupting the aggregates. Spectacular examples of these various structures are preserved in the Tuolumne Batholith, California. Detailed mechanisms for the formation of many of the structures are not well understood, though the formation of cumulates in vertical layers suggests that sorting and filter pressing during flow and resulting strain of crystal mushes may play important roles.

Sensitivity experiments with general circulation models show that uplift of plateau and mountain regions in Southern Asia and the American west during the late Cenozoic was an important factor in the evolution of Northern Hemisphere climate. The climatic trends simulated in the uplift experiments agree in direction with most trends observed in the geological record, including the tendencies toward greater regional differentiation of climate, and particularly the fragmentation into wetter and drier climatic patterns at middle latitudes. These climatic trends result from (1) increased orographic diversion of the mid-latitude westerlies, and (2) increased summer heating and winter cooling over the plateaus, which enhances seasonally reversing (monsoonal) changes in wind directions.

Most previous hypotheses addressing the physical impact of orography on climate have focused on mountain ranges and have stressed relatively local responses such as upslope precipitation maxima, cooling of mountain crests due to lapse-rate effects on rising terrain, and lee-side rainshadow effects. In contrast, our results emphasise the importance of large-scale plateau orography. By redirecting the basic directions of wind flow both at surface and upper-tropospheric levels, these rising plateaux cause far-reaching climatic changes that extend across the continents as well as over the oceans.

During recent years there has been a very great advance in our knowledge of the minute structure and function of the various regions and glands of the alimentary canal of the prosobranch and pulmonate gastropods, without any corresponding increase in our understanding of the opisthobranch and, in particular, of the nudibranch gut. That the emphasis should be laid on the streptoneurous and the land-living forms is obviously due to their abundance and to the fact that they include almost all the familiar types of gastropod mollusc, but it is disappointing that, at the moment of writing, there exists only one account of the histology and function of the digestive system of either a dorid or an æolid—to confine attention to the two main types of nudibranch molluscs which occur in British waters—that of Millott (1937b). This is perhaps more noticeable when it is recalled that the nudibranchs have specialised along distinctly unusual lines, many of which are intimately associated with the food and feeding habits which the animals have adopted. The æolids, it is true, have attracted a lively interest in connexion with their possession of nematocysts, but the majority of investigators have apparently been content to trace these into the cnidosacs without concerning themselves over the other constituents of the animal's food.

Using zircons taken from two granite plutons, Strontian (Caledonian, northwestern Scotland) and Kameruka (Bega Batholith, southeastern Australia), this study presents observations that have a bearing on refractory zircons as provenance indicators. Two broad textural types of refractory zircon were identified: (1) those which show simple two-stage growth histories; and (2) those which have apparently undergone repeated periods of growth, resorption, mechanical abrasion, fracturing and fracture-healing. SHRIMP U-Pb ages obtained from the Kameruka zircons indicate that the cores are the textural manifestation of inheritance. The shapes of refractory cores are not unambiguously indicative of their ultimate origin, since they may also be modified by processes that occur before and after incorporation into the magma. The cores within the two populations show a great diversity in types and styles of zoning, and in composition, implying that they have not chemically equilibrated internally, or externally with their host melt.

Evidence is given that many of the Mesozoic, geosynclinal serpentines of Turkey are deep-sea lavas, and that they functioned as efficient fertilizers of surface waters leading to rich crops of radiolaria now preserved in cherts and limestones. Furthermore, in anticlinals at Ankara and Alaca Höyük, Palæozoic, Triassic and Jurassic rocks in the condition of a tectonic mélange are overlain unconformably by Upper Cretaceous and Eocene; while, in associated synclinals at Ayas and Kirsehir, crystalline schists and granites, probably Palæozoic or earlier, emerge from beneath unconformable Oligocene. It is suggested that these crystallines are parts of a Pontic nappe which advanced southwards during Upper Cretaceous times, in part overriding and breaking local rocks to yield the Ankara Mélange, in part bulldozing them forwards to form the Taurus Mountains. The displacement along the Anatolian Thrust at the base of the invading Pontic nappe is reckoned at about 350 km.

A model for the early Palaeozoic metamorphic history of the Midland Valley and adjacent areas to the S in Scotland, England and Ireland is based on the results of new field mapping, thin section petrography, electron probe microanalysis, X-ray diffractometry, conodont and palynomorph colouration and graptolite reflectance measurement.

The oldest metamorphic rocks of the Midland Valley of Scotland, excluding xenoliths in post-Silurian lavas, are possibly the blueschist occurrences in the melange unit of the Ballantrae complex. These may be tectonised remnants of (?)pre-Arenig ocean-floor subducted during closure of the Iapetus Ocean. In the early Ordovician, the melange terrane was dynamothermally metamorphosed during obduction of newly-formed ocean crust. The obduction process piled up a thick sequence of various ocean-floor types such that burial metamorphism in parts reached pumpellyite-actinolite facies; elsewhere prehnite-pumpellyite and zeolite facies was attained.

Whilst the Midland Valley acted as an inter- or fore-arc basin during the Late Ordovician and Silurian and experienced burial metamorphism, an accretionary prism was formed to the S. Accretion, tectonic burial and metamorphism of ocean-floor and trench sediment was continuous in the Southern Uplands and the Longford-Down massif of Ireland through Late Ordovician to Late Silurian times. Rocks at the present-day surface vary from zeolite facies to prehnitepumpellyite facies. Silurian trench-slope basin sediments can be recognised in part by their lower grade of burial metamorphism. Greenschist facies rocks of the prism probably lie close to the surface.

The Lake District island-arc terrane of Northern England has an early Ordovician history of burial metamorphism up to prehnite-pumpellyite facies. The Late Ordovician and Silurian metamorphic history is one of sedimentary burial complicated by tectonism and intrusion of granite plutons to a relatively high level. The Iapetus suture is marked by a weak contrast in metamorphic grade.