In his “lost notebook,” Ramanujan used iterated derivatives of two theta functions to define sequences of q-series $\{U_{2t}(q)\}$ and $\{V_{2t}(q)\}$ that he claimed to be quasimodular. We give the first explicit proof of this claim by expressing them in terms of “partition Eisenstein series,” extensions of the classical Eisenstein series $E_{2k}(q),$ defined by $$ \begin{align*}\lambda=(1^{m_1}, 2^{m_2},\dots, n^{m_n}) \vdash n \ \ \ \longmapsto \ \ \ E_{\lambda}(q):= E_2(q)^{m_1} E_4(q)^{m_2}\cdots E_{2n}(q)^{m_n}. \end{align*} $$
For functions $\phi : \mathcal {P}\mapsto {\mathbb C}$ on partitions, the weight $2n$ partition Eisenstein trace is $$ \begin{align*}\operatorname{\mathrm{Tr}}_n(\phi;q):=\sum_{\lambda \vdash n} \phi(\lambda)E_{\lambda}(q). \end{align*} $$
For all t, we prove that $U_{2t}(q)=\operatorname {\mathrm {Tr}}_t(\phi _U;q)$ and $V_{2t}(q)=\operatorname {\mathrm {Tr}}_t(\phi _V;q),$ where $\phi _U$ and $\phi _V$ are natural partition weights, giving the first explicit quasimodular formulas for these series.