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How Music and Our Faculty for Music Are Made for Each Other

Published online by Cambridge University Press:  18 September 2024

Abstract

This study relies on the prevalence of certain structures that largely distinguish the creation and reception of music from that of language – namely, temporal grids, scalar grids, and segments with their repetitions – to construct a model of the human cognitive faculty for music that allows humans to make music the way they do. The study draws on research and thought in philosophy (including phenomenology), linguistics, psychology, and neurology, coupled with musicology, to produce a model of a human capacity to make complex comparisons between ongoing sound sequences and those simultaneously reconstructed from memory by registering the relativities within their flow. This model is then used in a consideration of how the faculty for music interacts with the faculty for language in the experience of song and a consideration of how a similar cognitive capacity for music might be identified in other species.

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Article
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© The Author(s), 2024. Published by Cambridge University Press on behalf of the Royal Musical Association

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Footnotes

I thank Jeremy Day-O’Connell, Gina Fatone, Mary Hunter, Daniel Leech-Wilkinson, Ralph Locke, and the reviewers for this journal, all of whom have given me valuable responses and suggestions.

References

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2 Languages of many other cultures use terms both broader and narrower than the European word music, rather than any term equivalent in its coverage, even if ethnomusicologists say that every culture has something that speakers of English might want to call music. See Bruno Nettl, ‘Music’, Grove Music Online, 2001 <www.oxfordmusiconline.com> (updated and revised 1 July 2014), II and II.7. But then, the European tradition itself was ambiguous from the start: Plato used the word mousikē to refer to the whole realm of the Muses, which included language and dance and other activities beyond what we call music, and yet he also wrote of it as the art (tekhnē) of modes and rhythms, the elements most defining of music in our narrow sense. Compare Plato’s language about the Muses in Phaedrus 259b–d to his language about harmonies and rhythms in Alcibiades I 108c and Republic 398c–400c.

3 See Patrick Savage and others, ‘Statistical Universals Reveal the Structures and Functions of Human Music’, Proceedings of the National Academy of Sciences USA, 112 (2015), 8987–92. The features of music described in my study have considerable overlap with those that Savage and colleagues propose.

4 See Stevens, Catherine J. and Byron, Tim, ‘Universals in Music Processing: Entrainment, Acquiring Expectations, and Learning’, in Oxford Handbook of Music Psychology, ed. Hallam, Susan and others, 2nd edn (Oxford University Press, 2016), 2032 Google Scholar. The processes described in my study have overlaps with those Stevens and Byron propose, especially in the realms of response to temporal structures and the processes of segmenting and grouping. The main differences lie in their concern with emotional response to generic structures, something I do not discuss, and their lack of recognition of the comparative processes that I identify as essential to the cognitive faculty for music.

5 Aristoxenus of Tarentum, Elementa Harmonica (Armonika Stoikheia), trans. Andrew Barker, in Barker, Greek Musical Writings, vol. 2: Harmonic and Acoustic Theory (Cambridge University Press, 1989), 15.

6 See the survey by Ian Cross and Elizabeth Tolbert, ‘Music and Meaning’, in Oxford Handbook of Music Psychology, ed. Hallam and others, 33–45.

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10 The music theory traditions of many cultures are primarily concerned with how to create good courses of musical events or how to track them.

11 We might suppose that Aristoxenus, as a pupil of Aristotle, at least understood the concept of memory that Aristotle articulated in the treatise known as De memoria et reminiscentia. On the implications of Aristotle’s ideas on memory for music, see Wiskus, Jessica, ‘On Music and Memory through Mnēmē and Anamnēsis ’, Research in Phenomenology 48 (2018), 346–64CrossRefGoogle Scholar.

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13 E. R. Clay (E. Robert Kelly), The Alternative: A Study in Psychology (Macmillan, 1882), 167. What Kelly called the ‘specious present’ the psychologist Daniel Stern later called the ‘present moment’, within which the ‘temporal contour’ of an experience (such as of a musical phrase) gives us the highly specific and subjective feeling that he calls a ‘vitality affect’. Daniel Stern, The Present Moment in Psychotherapy and Everyday Life (W. W. Norton, 2004), especially ch. 4.

14 William James, The Principles of Psychology, 2 vols (Henry Holt, 1890; repr. Dover Publications, 1950), vol. 1, ch. 15, pp. 609, 610.

15 The term echoic memory was introduced by Ulric Neisser in Cognitive Psychology (Appleton-Century-Crofts, 1967). Alan Baddeley describes the concept of auditory sensory memory in his standard textbook on memory, Essentials of Human Memory (Psychology Press, Taylor & Francis Group, 2014), 10–14.

16 James, Principles, vol. 1, ch. 9, pp. 240, 241.

17 Brentano, Franz, Psychology from an Empirical Standpoint, ed. Oskar Kraus, , English ed. Linda McAlister, trans. Antos Rancurello, D.B. Terrell, and Linda McAlister (Humanities Press, 1973), 168 Google Scholar. Original German edition, Leipzig: Duncker & Humblot, 1874. James, in a footnote to the passage cited here (p. 240) calls Brentano’s account ‘as good as anything with which I am acquainted’.

18 Husserl, Edmund, A Phenomenology of the Consciousness of Internal Time , trans. Brough, John, in The Essential Husserl: Basic Writings in Transcendental Phenomenology, ed. Donn Welton, (Indiana University Press, 1999)Google Scholar, section 2 (‘Analysis of the Consciousness of Time’), §10, 189. Section 2 is based on Husserl’s lectures in 1905.

19 Even though Aristoxenus was a noted disciple of Aristotle, it can hardly be surprising that these philosophers did not know or refer to this passage, since it occurs in a music treatise, one of the few texts by Aristoxenus to survive.

20 It is telling that in a century or more of phenomenological thinking about the experiencing of time, music has retained this exemplifying position. The phenomenological psychologist Daniel Stern, for example, in his study of The Present Moment in Psychotherapy and Everyday Life, 26, points to the musical phrase as ‘the musical analog of a present moment in ordinary life’: ‘it is felt to occur during a moment that is not instantaneous, but also not parceled out in time into sequential bits like the written notes’.

21 Huron, David, ‘A Psychological Approach to Musical Form: The Habituation-Fluency Theory of Repetition’, Current Musicology, 96 (2013), 7.Google Scholar

22 Ulric Neisser, Cognition and Reality: Principles and Implications of Cognitive Psychology (W. H. Freeman, 1976), 20–24.

23 George Mashour and others, ‘Conscious Processing and the Global Neuronal Workspace Hypothesis’, Neuron, 105 (2020), 782.

24 Alan Baddeley, for instance, titles the ninth chapter of his Essentials of Human Memory ‘Retrieval’.

25 Damasio, Antonio, Self Comes to Mind: Constructing the Conscious Brain (Pantheon Books, 2010), 133.Google Scholar

26 Ibid., 141–42.

27 Edelman, Gerald, Second Nature: Brain Science and Human Knowledge (Yale University Press, 2006), 28.Google Scholar

28 Bob Snyder, Music and Memory: An Introduction (MIT Press, 2000). See especially chs 4 (Short-Term and Working Memory) and 6 (Long-Term Memory).

29 Turino, Thomas, Music as Social Life: The Politics of Participation (University of Chicago Press, 2008), 18.Google Scholar

30 Among recent studies of the evolution of music, a few that focus particularly on the evolution of the faculty, or capacity, for music are Mithen, Steven, The Singing Neanderthals: The Origins of Music, Language, Mind and Body (Harvard University Press, 2006)Google Scholar; Tomlinson, Gary, A Million Years of Music: The Emergence of Human Modernity (Zone Books, 2015)CrossRefGoogle Scholar; and Henkjan Honing, ed., The Origins of Musicality (MIT Press, 2018).

31 Plato, Philebus, 17d, trans. R. Hackforth (Cambridge University Press, 1945), repr. in The Collected Dialogues of Plato, ed. Edith Hamilton and Huntington Cairns (Princeton University Press, 1961), 1093.

32 Measuring musical rhythms from the onset of one sound-event to the onset of the next is a standard technique in studies of music in both humans and other species. See, for instance, De Gregorio, and others, ‘Categorical Rhythms in a Singing Primate’, Current Biology, 31 (October 25, 2021), R1379 CrossRefGoogle Scholar. Sarah Hawkins explains why the ‘p-centres’ (psychological moments of occurrence) of musical notes are more closely aligned with their onsets than is the case with the p-centres of syllables in spoken words, in ‘Situational Influences on Rhythmicity in Speech, Music, and Their Interaction’, Philosophical Transactions of the Royal Society B, 369 (2014), 20130398, 3. Dancers, of course, characteristically move to the onsets of musical sounds.

33 The term categorical rhythm for small-integer rhythmic relationships seems to be favoured especially in biologists’ studies of non-human species. See, for instance, Roeske, Tina and others, ‘Categorical Rhythms Are Shared between Songbirds and Humans’, Current Biology, 30 (21 September 2020), 3544–55.CrossRefGoogle Scholar

34 On non-equal beats, see London, Justin, Hearing in Time: Psychological Aspects of Musical Meter (Oxford University Press, 2012), ch. 8.CrossRefGoogle Scholar

35 On Plato’s role in forming the Greek concept of rhythm, see Benveniste, Émile, ‘The Notion of “Rhythm” in Its Linguistic Expression’, Problems in General Linguistics, trans. Mary Elisabeth Meek (University of Miami Press, 1971), 281–88Google Scholar. Originally published in French in 1951. In connecting musical rhythms to ‘the performer’s bodily movements’, Plato can be considered a forerunner of a wide range of modern concepts, from Mark Johnson’s idea that music’s meaning is embodied (see his Meaning of the Body) and Arnie Cox’s exploration of mimetic and metaphorical modelling of music on human bodily experience in Music and Embodied Cognition: Listening, Moving, Feeling, and Thinking (Indiana University Press, 2016) to psychological theories of embodied musical cognition, as in Schiavio, Andrea and others, ‘Music in the Flesh: Embodied Simulation in Musical Understanding’, Psychomusicology: Music, Mind, and Brain, 24.4 (2014), 340–43CrossRefGoogle Scholar.

36 London, Hearing in Time, 24.

37 Edward Large, ‘Resonating to Musical Rhythm: Theory and Experiment’, in Psychology of Time, ed. Simon Grondin (Emerald Group, 2008), 189–232, offers a ‘neural resonance theory’ of how listeners ‘experience dynamic temporal patterns, and hear musical events in relation to these patterns, because they are intrinsic to the physics of the neural systems involved in perceiving, attending, and responding to auditory stimuli’.

38 Jacoby, Nori and McDermott, Josh, ‘Integer Ratio Priors on Musical Rhythm Revealed Cross-Culturally by Iterated Reproduction’, Current Biology, 27 (2017), 359–70CrossRefGoogle ScholarPubMed.

39 The scheme proposed here agrees in large part with the ‘separable components’ that Sonja Kotz and colleagues specify as ‘underlying rhythm cognition’. What I am calling ‘abstracting the grid’, for example, they call ‘beat extraction from complex auditory patterns’. But their scheme lacks the prior stage I describe as ‘accumulating the sounds into measured rhythmic “figures”’. See S. A. Kotz and others, ‘The Evolution of Rhythm Processing’, Trends in Cognitive Sciences, 22.10 (October 2018), 896–97. Edward Large by contrast does account for the ‘pulse and meter’ to emerge from the complex sounding sequence of note-lengths through a process he calls induction; at the same time, he considers pulse and metre to be ‘intrinsic to the physics of neural oscillation. All that is then required is coupling to a rhythmic stimulus.’ Large, ‘Resonating to Musical Rhythm’, 193, 223.

40 These effects have been explored in studies including McNeill, William, Keeping Together in Time: Dance and Drill in Human History (Harvard University Press, 1995)Google Scholar; Turino, Music as Social Life; Hove, Michael and Risen, Jane, ‘It’s All in the Timing: Interpersonal Synchrony Increases Affiliation’, Social Cognition, 27.6 (2009), 949–61CrossRefGoogle Scholar; and Cross, Liam and others, ‘How Moving Together Binds Us Together: The Social Consequences of Interpersonal Entrainment and Group Processes’, Open Psychology, 1 (2019), 273302 CrossRefGoogle Scholar.

41 The study of musical entrainment was greatly advanced by the publication of Martin Clayton and others’ ‘In Time with the Music: The Concept of Entrainment and its Significance for Ethnomusicology’, European Meetings in Ethnomusicology, 11 (2005), ESEM Counterpoint 1, 1–82, now on Durham Research Online: https://dro.dur.ac.uk/8713/. The present study benefits, for instance, from these authors’ recognition (p. 15) that ‘the motor system is not only responsible for producing a rhythm, but is also involved in the perception of rhythm’. A similar recognition of the role of memory has been less apparent so far in the work of these and other authors investigating musical entrainment.

42 Keil, Charles, ‘Participatory Discrepancies and the Power of Music’, Cultural Anthropology, 2.3 (August 1987), 275–83CrossRefGoogle Scholar.

43 Kotz and others, ‘Evolution of Rhythm Processing’, 900.

44 Ibid., 901.

45 Clayton and others, ‘In Time with the Music’, 20–25.

46 See Widdess, Richard, ‘Involving the Performers in Transcription and Analysis: A Collaborative Approach to Dhrupad’, Ethnomusicology, 38.1 (1994), 5979 CrossRefGoogle Scholar, especially pp. 65–68.

47 Frank Kouwenhoven, ‘Some Remarks on Music as Reorganized Time’, Commentary in Clayton et al., ‘In Time with the Music’, 88–92.

48 Victor Zuckerkandl writes: ‘Music is temporal art in the special sense that in it time reveals itself to experience’. Sound and Symbol: Music and the External World, trans. Willard Trask (Pantheon, 1956), 200.

49 The use of discrete pitches counts as one of the most universal of musical features among those considered in Savage and others, ‘Statistical Universals’.

50 Trehub, Sandra, ‘Human Processing Predispositions and Musical Universals’, in The Origins of Music, ed. Wallin, Nils and others (MIT Press, 2000), 427–48Google Scholar, esp. pp. 428–31.

51 Alexander Ellis, ‘On the Musical Scales of Various Nations’, Journal of the Society of Arts, 33 (1885), 526. Reprinted in Kay Kaufman Shelemay, ed., Garland Library of Readings in Ethnomusicology 7 (Garland Publishing, 1990), 1–43. This observation places Ellis in Aristoxenus’ rather than Pythagoras’ line of scale theorists.

52 Even infants evidently process music more effectively when it is made of uneven-step rather than even-step scales. See Sandra Trehub and others, ‘Infants’ and Adults’ Perception of Scale Structure’, Journal of Experimental Psychology: Human Perception and Performance, 25.4 (1999), 965–75.

53 It is precisely for disorienting effects that musicians sometimes adopt equal-step scales (e.g., the chromatic and whole-tone scales in Western music).

54 By ingenious experimental design Sandra Trehub and colleagues produced evidence that six- to eight-month-old infants can respond differentially to a six-note melody when it is repeated with a single pitch significantly changed. See Sandra Trehub and others, ‘Infants’ Perception of Melodies: Changes in a Single Tone’, Infant Behavior and Development, 8 (1985), 213–23.

55 Listeners frequently find music at certain moments hair-raising, or chilling, or tear-inducing, and studies have explored what kinds of events in music set off such physical responses. See Donald Hodges, ‘Bodily Responses’, Oxford Handbook of Music Psychology, ed. Hallam and others, 183–96.

56 Jan Gorisch and others, ‘Pitch Contour Matching and Interactional Alignment across Turns: An Acoustic Investigation’, Language and Speech, 55.1 (2012), 57–76; Juan Pablo Robledo and others, ‘Pitch-Interval Analysis of “Periodic” and “Aperiodic” Question+Answer Pairs’, Speech Prosody 2016, 1071–75. https://eprints.whiterose.ac.uk/96973/.

57 Ladd, D. Robert, Intonational Phonology, 2nd edn (Cambridge University Press, 2008), 196 CrossRefGoogle Scholar. Likewise Ray Jackendoff writes that ‘there is no convincing analogue in language to the music use of pitch space, despite their making use of the same motor capacities in the vocal tract’. ‘Parallels and Nonparallels between Language and Music’, Music Perception, 26.3 (2009), 200. For further comparison of language to music as a sound system, see Patel, Aniruddh, Music, Language and the Brain (Oxford University Press, 2008)Google Scholar, especially ch. 2.3, ‘Linguistic Sound Systems’.

58 Roger Chaffin and others, ‘Performing from Memory’, Oxford Handbook of Music Psychology, ed. Hallam and others, 560.

59 Baddeley, Essentials, 21–25.

60 Bob Snyder, ‘Memory for Music’, Oxford Handbook of Music Psychology, ed. Hallam and others, 168.

61 Margulis, Elizabeth Hellmuth, On Repeat: How Music Plays the Mind (Oxford University Press, 2014), 23.Google Scholar

62 As Nicolas Ruwet writes, ‘if it is true that variation is the soul of all music […] it is no less true that to say variation is to say repetition: there can only be variation on a given level, whatever that may be, if there is at the same time repetition on another level’. Langage, Musique, Poésie (Editions du Seuil, 1972), 136.

63 Huron, David, Sweet Anticipation: Music and the Psychology of Expectation (MIT Press, 2006).CrossRefGoogle Scholar

64 V. Salimpoor and others, ‘Anatomically Distinct Dopamine Release during Anticipation and Experience of Peak Emotion to Music’, Nature Neuroscience, 14.2 (February 2011), 257–64. For a survey of research on the roles of neurotransmitters in musical experiences, see Koshimori, Yuko, ‘Neurochemical Responses to Music’, in Oxford Handbook of Music and the Brain, ed. Hodges, Donald and Thaut, Michael (Oxford University Press, 2019), ch. 14.Google Scholar

65 Margulis, On Repeat, 5.

66 Aristotle, Poetics, 1448b, trans. W. H. Fyfe in Aristotle in 23 Volumes, vol. 23 (Harvard University Press, 1932).

67 John Booth Davies described what he called the ‘Darling, They’re Playing Our Tune’ phenomenon as a distinctive trait of musical experience in The Psychology of Music (Stanford University Press, 1978).

68 See Gabrielsson, Alf, Strong Experiences with Music: Music Is Much More Than Just Music, trans. Bradbury, Rod (Oxford University Press, 2011)Google Scholar; Herbert, Ruth, Everyday Music Listening: Absorption, Dissociation, and Trancing (Routledge, 2012).Google Scholar

69 See Gilbert Rouget, Music and Trance: A Theory of the Relations between Music and Possession, trans. author, rev. Brunhilde Biebuyck (University of Chicago Press, 1985; original French edn 1980), the classic study of the subject. Judith Becker’s study, Deep Listeners: Music, Emotion, and Trancing (Indiana University Press, 2004), brings more recent neuroscientific findings to bear on the subject.

70 The model by Chomsky and associates appears in Hauser, Marc, Chomsky, Noam, and Fitch, W. Tecumseh, ‘The Faculty of Language: What Is It, Who Has It, and How Did It Evolve?Science, 298 (2002), 1569–79CrossRefGoogle ScholarPubMed, updated in Berwick, Robert C. and Chomsky, Noam, ‘The Biolinguistic Program: The Current State of Its Development’, The Biolinguistic Enterprise: New Perspectives on the Evolution and Nature of the Human Language Faculty, ed. Di Sciullo, Anna-Maria and Boeckx, Cedric (Oxford University Press, 2011), 1941 Google Scholar. The Jackendoff model appears in Jackendoff, Ray, Foundations of Language: Brain, Meaning, Grammar, Evolution (Oxford University Press, 2002)CrossRefGoogle Scholar and Jackendoff, ‘What Is the Human Language Faculty? Two Views’, Language, 87.3 (2011), 586–624. See also the exchange between the two parties in Pinker, Steven and Jackendoff, Ray, ‘The Faculty of Language: What’s Special about It?Cognition, 95 (2005), 201–36CrossRefGoogle Scholar; Fitch, W. Tecumseh, Hauser, Marc, and Chomsky, Noam, ‘The Evolution of the Language Faculty: Clarifications and Implications’, Cognition, 97 (2005), 179210 CrossRefGoogle ScholarPubMed; Jackendoff, Ray and Pinker, Steven, ‘The Nature of the Language Faculty and Its Implications for Evolution of Language’, Cognition, 97 (2005), 211–25CrossRefGoogle Scholar.

71 Hauser, Chomsky, and Fitch, in ‘The Faculty of Language’, 1571, claim that recursion is a ‘core property’ of the language faculty and ‘appears’ possibly ‘to lack any analog in […] other domains as well’. Jackendoff, in ‘What Is the Human Language Faculty’, 591–99, claims that recursion is part of a general human capacity that includes music.

72 Jackendoff, ‘What Is the Human Language Faculty’, 599. On p. 587, n. 3, he notes that Hauser and others, in ‘The Faculty of Language’, p. 1571, ‘specifically exclude memory from [the language faculty, in the broad sense], for reasons unclear to me’.

73 The vast lexicon of words in any language is of course built from a much smaller set of phonemes.

74 See Cuddy, Lola and others, ‘Memory for Melodies and Lyrics in Alzheimer’s Disease’, Music Perception: An Interdisciplinary Journal, 29.5 (2012), 479–91CrossRefGoogle Scholar.

75 Booth, Mark C., The Experience of Song (Yale University Press, 1981), 78.Google Scholar

76 Charles Dickens, in A Christmas Carol (1843), cited the song as ‘God bless you, merry Gentlemen’.

77 Wikipedia, ‘God Rest You Merry, Gentlemen’, consulted 1 September 2022.

78 Damasio, Self Comes to Mind, 211.

79 See Salimpoor and others, note 64 above.

80 These phrases can be heard at 0:53 sung by Ella Fitzgerald in The Rodgers and Hart Song Book <https://www.youtube.com/watch?v=t78xp_BHXlA> (accessed 9 May 2024).

81 Lucretius, The Nature of Things [De rerum natura], Book 5, ll. 1379ff., trans. A. E. Stallings (Penguin Books, 2007), 192.

82 Martinelli, Dario, Of Birds,Whales, and Other Musicians: An Introduction to Zoomusicology (University of Scranton Press, 2009), 217.Google Scholar

83 See Sayigh, Laela and Janik, Vincent, ‘Cetacean Communication’, in Deep Thinkers: Inside the Minds of Whales, Dolphins, and Porpoises, ed. Janet Mann, (University of Chicago Press, 2017), 7276 Google Scholar.

84 Thorpe, W. H., Bird-Song: The Biology of Vocal Communication and Expression in Birds (Cambridge University Press, 1961), 15.Google Scholar

85 Roger Payne and Scott McVay, ‘Songs of Humpback Whales’, Science, 173/3997 (13 August 1971), 585–97.

86 Katharine Payne, ‘The Progressively Changing Songs of Humpback Whales: A Window on the Creative Process in a Wild Animal’, in The Origins of Music, ed. Wallin and others, 139.

87 Danielle Cholewiak and others, ‘Humpback Whale Song Hierarchical Structure: Historical Context and Discussion of Current Classification Issues’, Marine Mammal Science, 29.3 (July 2013), E312–E332, E314.

88 Mercado, Eduardo III and Handel, Stephen, ‘Understanding the Structure of Humpback Whale Songs’, Journal of the Acoustical Society of America, 132 (2012), 2947–50CrossRefGoogle ScholarPubMed.

89 Katharine Payne and Roger Payne, ‘Large Scale Changes over 19 Years in Songs of Humpback Whales in Bermuda’, Zeitschrift für Tierpsychologie (now Ethology), 68 (1985), 91.

90 Elemans, and others, ‘Evolutionary Novelties Underlie Sound Production in Baleen Whales’, Nature, 21 (2024)Google Scholar, Discussion. See also Adam, Olivier and others, ‘New Acoustic Model for Humpback Whale Sound Production’, Applied Acoustics, 74 (2013), 1182–90CrossRefGoogle Scholar.

91 Elemans and others, ‘Evolutionary Novelties’.

92 Personal communication, Hansen Johnson, Dalhousie University, 14 March 2016.

93 Payne, ‘The Progressively Changing Songs’, 138. These group alterations were first described in Katharine Payne and others, ‘Progressive Changes in the Songs of Humpback Whales (Megaptera novaeangliae): A Detailed Analysis of Two Seasons in Hawaii’, in Communication and Behavior of Whales, ed. Roger Payne (Westview Press, 1983), 9–57.

94 Further research has shown that this ‘evolution’, in which the content of a song grows steadily more complex, is supplanted at times by ‘revolutionary’ events, in which songs are ‘always completely replaced with a simpler song’. Allen, Jenny and others, ‘Cultural Revolutions Reduce Complexity in the Songs of Humpback Whales’, Proceedings of the Royal Society B, 285 (2018), 3.Google ScholarPubMed

95 Linda Guinee and Katharine Payne, ‘Rhyme-like Repetitions in Songs of Humpback Whales’, Ethology, 79 (1988), 295–306. See also Payne and others, ‘Progressive Changes in the Songs’.

96 Herman, Louis, ‘The Multiple Functions of Male Song within the Humpback Whale (Megaptera novaeangliae) Mating System: Review, Evaluation, and Synthesis’, Biological Reviews of the Cambridge Philosophical Society, 92 (2017), 1795–818CrossRefGoogle ScholarPubMed. The situation is described succinctly – ‘While song is well described, its function is not yet fully understood’ – by Dana Cusano and others in ‘Socially Complex Breeding Interactions in Humpback Whales Are Mediated Using a Complex Acoustic Repertoire’, Frontiers in Marine Science, https://doi.org/10.3389/fmars.2021.665186.

97 A kind of comparison across time is found even in bacteria, in which, according to Peter Godfrey-Smith, ‘one mechanism registers what conditions are like right now, and another records how things were a few moments ago’. Godfrey-Smith, Peter, Other Minds: The Octopus, the Sea, and the Deep Origins of Consciousness (Farrar, Straus and Giroux, 2016), 17 Google Scholar. But that capacity does not involve anything like the comparison of simultaneously remembered and perceived sound sequences required in musical processing.