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New data on Homocladus grandis, a Permian stem-mantodean (Polyneoptera: Dictyoptera)

Published online by Cambridge University Press:  20 May 2016

Olivier Béthoux ,
Roy J. Beckemeyer ,
Michael S. Engel  and
Joseph D. Hall
Olivier Béthoux
Affiliation:
Freiberg University of Mining and Technology, Institute of Geology, Department of Palaeontology, Bernhard-von-Cotta Str. 2, D-09596 Freiberg, Germany; Senckenberg Natural History Collections of Dresden, Museum of Zoology, Königsbrücker Landstraße 159, D-01109 Dresden, Germany,
Roy J. Beckemeyer
Affiliation:
Division of Entomology (Paleoentomology), Natural History Museum, 1501 Crestline Drive—Suite 140, University of Kansas, Lawrence, Kansas 66049-2811;
Michael S. Engel
Affiliation:
Division of Entomology (Paleoentomology), Natural History Museum, and Department of Ecology and Evolutionary Biology, 1501 Crestline Drive—Suite 140, University of Kansas, Lawrence, Kansas 66049-2811 Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024-5192,
Joseph D. Hall
Affiliation:
Chaparral Energy, 701 Cedar Lake Boulevard, Oklahoma City, OK 73114
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Abstract

Representatives of the family Strephocladidae have been considered as fossil relatives (i.e., stem-group) of Mantodea (mantises) based on characters of the forewing morphology. Here we describe new specimens from the Wellington Formation that we assign to the strephocladid species Homocladus grandis Carpenter, 1966. The range of morphological variation exhibited by the new material, in addition to wing morphology variability documented in extant mantises and roaches, suggest that H. ornatus Carpenter, 1966 and Paracladus retardatus Carpenter, 1966, reported from the same formation, are new junior subjective synonyms of H. grandis. We describe the first hind wing for this species based on a well-preserved specimen. It exhibits a combination of character states unique to dictyopteran insects.

Type
Research Article
Information
Journal of Paleontology , Volume 84 , Issue 4 , July 2010 , pp. 746 - 753
Copyright
Copyright © The Paleontological Society

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