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Late Pleistocene Bison antiquus from Orcas Island, Washington, and the biogeographic importance of an early postglacial land mammal dispersal corridor from the mainland to Vancouver Island

Published online by Cambridge University Press:  20 January 2017

Michael C. Wilson ,
Stephen M. Kenady  and
Randall F. Schalk
Michael C. Wilson*
Affiliation:
Department of Earth and Environmental Sciences, Douglas College, P.O. Box 2503, New Westminster, BC, Canada V3L 5B2 Department of Archaeology, Simon Fraser University, Burnaby, BC Canada V5A 1S6
Stephen M. Kenady
Affiliation:
Cultural Resource Management, 5319 Cedar Ridge Place, Sedro-Woolley, WA 98284, USA
Randall F. Schalk
Affiliation:
Cascadia Archaeology, P.O. Box 51058, Seattle, WA 98115-1058, USA
*
*Corresponding author. Email Address:[email protected] (M.C. Wilson).
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Abstract

A Bison antiquus cranium and partial skeleton from Ayer Pond wetland on Orcas Island, San Juan Islands, Washington, date to 11,760 ± 70 14C yr BP. They lay in lacustrine sediments below peat, unconformably above emergent Everson Glaciomarine Drift (> 12,000 14C yr BP). Several bison finds in similar contexts on Orcas and Vancouver Islands dating between 11,750 and 10,800 14C yr BP indicate an early postglacial land mammal dispersal corridor with reduced water barriers between mainland and islands. New bison dates and published shell dates allow estimation of early postglacial relative sea-level trends for the San Juans, with a drop below modern datum ∼ 12,000 14C yr BP, and assist in evaluation of marine reservoir corrections. Emergence by ∼ 60 m is suggested by data from nearby areas. A tundra-like or meadow community and succeeding open pine parkland before 11,000 14C yr BP supported bison but horn-core reduction suggests suboptimal forage or restricted habitat. Expanding mixed-conifer forests after 11,000 14C yr BP contributed to bison extirpation. Dispersing ungulates such as bison must have influenced island vegetation establishment and early succession. Possible evidence for butchering by early coastal people adds significance to the Ayer Pond discovery, given its pre-Clovis age.

Keywords

PaleogeographyPaleoenvironmentsBison antiquusDispersalPacific NorthwestIsostasySea levelPaleontologyArchaeologyMarine reservoir correction
Type
Articles
Information
Quaternary Research , Volume 71 , Issue 1 , January 2009 , pp. 49 - 61
Copyright
University of Washington

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References

Al-Suwaidi, M., Ward, B.C., Wilson, M.C., Hebda, R.J., Nagorsen, D.W., Marshall, D., Ghaleb, B., Wigen, R.J., and Enkin, R.J. Late Wisconsinan Port Eliza Cave deposits and their implications for human coastal migration, Vancouver Island, Canada. Geoarchaeology, An International Journal 21, (2006). 305–399.Google Scholar
Alley, N.F., and Chatwin, S.C. Late Pleistocene history and geomorphology, southwestern Vancouver Island, British Columbia. Canadian Journal of Earth Sciences 16, (1979). 1645–1657.Google Scholar
Anundsen, K., Abella, S., Leopold, E., Stuiver, M., and Turner, S. Late-Glacial and early Holocene sea-level fluctuations in the central Puget Lowland, Washington, inferred from lake sediments. Quaternary Research 42, (1994). 149–161.CrossRefGoogle Scholar
Armstrong, J.E., Crandell, D.R., Easterbrook, D.J., and Noble, J.B. Late Pleistocene stratigraphy and chronology in southwestern British Columbia and northwestern Washington. Geological Society of America Bulletin 65, (1965). 349–364.Google Scholar
Balzarini, M.A. Paleoecology of late Pleistocene glacial-marine sediments in northwestern Washington and southwestern British Columbia. Molnia, B.F. Glacial-marine Sedimentation. (1983). Plenum, New York. 571–592.Google Scholar
Bard, E. Geochemical and geophysical implications of the radiocarbon calibration. Geochimica et Cosmochimica Acta 62, (1998). 2025–2038.Google Scholar
Barnosky, C.W., Anderson, P.M., and Bartlein, P.J. The northwestern U.S. during deglaciation; vegetational history and paleoclimatic implications. Ruddiman, W.F., Wright, H.E. Jr. North America and Adjacent Oceans During the Last Deglaciation. Geological Society of America, The Geology of North America, vol. K-3. (1987). Boulder, CO, 289–321.Google Scholar
Barrie, J.V., and Conway, K.W. A Preliminary Interpretation of Surficial Marine Geology of Central and Northern Strait of Georgia, British Columbia. (2000). Geological Survey of Canada Current Research, 2000–A21, 7 ppGoogle Scholar
Barrie, J.V., and Hill, P.R. Holocene faulting on a tectonic margin: Georgia Basin, British Columbia, Canada. Geo-Marine Letters 24, (2004). 86–96.CrossRefGoogle Scholar
Barrie, J.V., Hill, P.R., Conway, K.W., Iwanowska, K., and Picard, K. Environmental Marine Geoscience 4. Georgia Basin: seabed features and marine geohazards. Geoscience Canada 32, (2005). 145–156.Google Scholar
Barton, B.R. Some notable finds of Columbian mammoths from Washington State. Washington Geology 27, (1999). 23–27.Google Scholar
Barton, B.R., Kester, P.R., (2001). Schreger angle analysis of the Kirkland/Lakeview Elementary School tusk: mammoth or mastodon?. [abstract] In: Storer, J.E. (Ed.), Canadian Quaternary Association Meetings, 2001, Program and Abstracts. Yukon Heritage Branch, Whitehorse, Yukon., Occasional Papers in Earth Sciences 1, p. 23.Google Scholar
Bedwell, S.F. Fort Rock Basin Prehistory and Environment. (1973). University of Oregon Press, Eugene, OR.Google Scholar
Bement, L.C. Bison Hunting at Cooper Site: Where Lightning Bolts Drew Thundering Herds. (1999). University of Oklahoma Press, Norman, OK. 227 ppGoogle Scholar
Blais-Stevens, A., Bornhold, B.D., Kemp, A.E.S., Dean, J.M., and Vaan, A.A. Overview of Late Quaternary stratigraphy in Saanich Inlet, British Columbia: results of Ocean Drilling Program Leg 169S. Marine Geology 174, (2001). 3–20.Google Scholar
Blais-Stevens, A., Clague, J.J., Mathewes, R.W., Hebda, R.J., and Bornhold, B.D. Record of large, Late Pleistocene outburst floods preserved in Saanich Inlet sediments, Vancouver Island, Canada. Quaternary Science Reviews 22, (2003). 2327–2334.CrossRefGoogle Scholar
Booth, D.B. Timing and processes of deglaciation along the southern margin of the Cordilleran ice sheet. Ruddiman, W.F., Wright, H.E. Jr. North America and Adjacent Oceans During the Last Deglaciation. Geological Society of America, The Geology of North America, vol. K-3. (1987). Boulder, CO, 71–90.Google Scholar
Booth, D.B. Glaciofluvial infilling and scour of the Puget Lowland, Washington, during ice-sheet glaciation. Geology 22, (1994). 695–698.Google Scholar
Booth, D.B., Troost, K.G., Clague, J.J., and Waitt, R.B. The Cordilleran ice sheet. Developments in Quaternary Science 1, (2003). 17–43.Google Scholar
Briggs, J.M., Knapp, A.K., and Brock, B.L. Expansion of woody plants in tallgrass prairie: a fifteen-year study of fire and fire-grazing interactions. American Midland Naturalist 147, (2002). 287–294.CrossRefGoogle Scholar
Brown, J.H., and Lomolino, M.V. Concluding remarks: historical perspective and the future of island biogeography. Global Ecology and Biogeography 9, (2000). 87–92.Google Scholar
Brown, K.J., and Hebda, R.J. Origin, development, and dynamics of coastal temperate conifer rainforests of southern Vancouver Island, Canada. Canadian Journal of Forestry Research 32, (2002). 353–372.CrossRefGoogle Scholar
Burns, K.C. A multi-scale test for dispersal filters in an island plant community. Ecography 28, (2005). 552–560.Google Scholar
Campbell, C., Campbell, I.D., Blyth, C.B., and McAndrews, J.H. Bison extirpation may have caused aspen expansion in western Canada. Ecography 17, (1994). 360–362.CrossRefGoogle Scholar
Canadian Hydrographic Service, (2005). Juan de Fuca Strait to Strait of Georgia. Nautical Chart, Scale 1:80 000.Google Scholar
Carlson, R.L. Cultural antecedents. Suttles, W. Handbook of North American Indians Vol. 7, (1990). Northwest Coast. Smithsonian Institution, Washington, DC. 60–69.Google Scholar
Chandler, A.C. A study of the skull and dentition of Bison antiquus Leidy, with special reference to material from the Pacific Coast. University of California Publications, Bulletin of the Department of Geology 9, (1916). 121–135.Google Scholar
Clague, J.J. Quadra Sand and its relation to the late Wisconsin glaciation of southwest British Columbia. Canadian Journal of Earth Sciences 13, (1976). 805–815.Google Scholar
Clague, J.J. Late Quaternary Geology and Geochronology of British Columbia, Part 1: Radiocarbon Dates. (1980). Geological Survey of Canada Paper 80-13, Ottawa, ON. 28 pp.Google Scholar
Clague, J.J. Late Quaternary Geology and Geochronology of British Columbia, Part 2: Summary and Discussion of Radiocarbon-dated Quaternary History. (1981). Geological Survey of Canada Paper 80-35, Ottawa, ON. 41 ppGoogle Scholar
Clague, J.J. Glacio-isostatic effects of the Cordilleran ice sheet, British Columbia, Canada. Smith, D.E., and Lawson, A.G. Shorelines and Isostasy. (1983). Academic Press, San Diego, CA. 321–343.Google Scholar
Clague, J.J., Harper, J.R., Hebda, R.J., and Howes, D.E. Late Quaternary sea-level and crustal movements, coastal British Columbia. Canadian Journal of Earth Sciences 19, (1982). 597–618.CrossRefGoogle Scholar
Clague, J.J., Mathewes, R.W., Guilbault, J.-P., Hutchinson, I., and Ricketts, B.D. Pre-Younger Dryas resurgence of the southwestern margin of the Cordilleran ice sheet, British Columbia, Canada. Boreas 26, (1997). 261–276.Google Scholar
Clague, J.J., and James, T.S. History and isostatic effects of the last ice sheet in southern British Columbia. Quaternary Science Reviews 21, (2002). 71–87.Google Scholar
Conroy, C.J., Demboski, J.R., and Cook, J.A. Mammalian biogeography of the Alexander Archipelago of Alaska: a north temperate nested fauna. Journal of Biogeography 26, (1999). 343–352.Google Scholar
Constible, J.M., Sweitzer, R.A., Van Vuren, D.H., Schuyler, P.T., and Knapp, D.A. Dispersal of non-native plants by introduced bison in an island ecosystem. Biological Invasions 7, (2005). 699–709.CrossRefGoogle Scholar
Conway, K.W., Barrie, J.V., and Hebda, R.J. Evidence for a Late Quaternary outburst flood event in the Georgia Basin, British Columbia. (2001). Geological Survey of Canada Current Research 2001-A13, Ottawa, ON. 15 Google Scholar
Coppedge, B.R., Leslie, D.M. Jr., and Shaw, J.H. Botanical composition of bison diets on tallgrass prairie in Oklahoma. Journal of Range Management 51, (1998). 379–382.Google Scholar
Croft, D.A., Heaney, L.R., Flynn, J.J., and Bautista, A.P. Fossil remains of a new, diminutive Bubalus (Artiodactyla: Bovidae: Bovini) from Cebu Island, Philippines. Journal of Mammalogy 87, (2006). 1037–1051.Google Scholar
Darwin, C. On the Origin of Species by Means of Natural Selection. (1859). First ed. John Murray, London.Google Scholar
Dethier, D.P., Pessl, F. Jr., Keuler, R.F., Balzarini, M.A., and Pevear, D.R. Late Wisconsinan glaciomarine deposition and isostatic rebound, northern Puget Lowland, Washington. Geological Society of America Bulletin 107, (1995). 1288–1303.Google Scholar
Dethier, D.P., Crane, W.H., Hirshfeld, M.A., Holbert, J.K., Morgan, W.S., Sylvester, R.E., (1996a). Morphology and seismic stratigraphy of marine banks, northern Puget Lowland, Washington. Geological Society of America Abstracts with Programs 28, A-435.Google Scholar
Dethier, D.P., White, D.P., Brookfield, C.M., (1996b). Maps of the Surficial Geology and Depth to Bedrock of False Bay, Friday Harbor, Richardson, and Shaw Island 7.5-minute Quadrangles, San Juan County, Washington. Washington State Department of Natural Resources, Division of Geology and Earth Resources, Open File Report 96-7, 7 pp.Google Scholar
Dethier, D.P., Nelson, R.E., Schildgen, T.F., (1998). Latest Pleistocene emergence and repopulation history at three sites in the northern Puget Lowland, Washington. Geological Society of America Abstracts with Programs 30, A-386.Google Scholar
Dixon, E.J. Human colonization of the Americas: timing, technology and process. Quaternary Science Reviews 20, (2001). 277–299.Google Scholar
Dorn, T.F., Fairhall, A.W., Schell, W.R., and Takashima, Y. Radiocarbon dating at the University of Washington I. Radiocarbon 4, (1962). 1–12.Google Scholar
Dyck, W., Lowdon, J.A., Fyles, J.G., Blake, W. Jr. Geological Survey of Canada radiocarbon dates V. Radiocarbon 8, (1966). 96–127.Google Scholar
Dyke, A.S., Dale, J.E., and McNeely, R.N. Marine mollusks as indicators of environmental change in glaciated North America during the last 18,000 years. Géographie physique et Quaternaire 50, (1996). 125–184.Google Scholar
Easterbrook, D.J. Late Pleistocene glacial events and relative sea-level changes in the northern Puget Lowland, Washington. Geological Society of America Bulletin 74, (1963). 1465–1484.Google Scholar
Easterbrook, D.J. Radiocarbon chronology of Late Pleistocene deposits in northwest Washington. Science 152, (1966). 764–767.Google Scholar
Easterbrook, D.J. Pleistocene chronology of the Puget Lowland and San Juan Islands, Washington. Geological Society of America Bulletin 80, (1969). 2273–2286.Google Scholar
Easterbrook, D.J., (1986). Stratigraphy and chronology of Quaternary deposits of the Puget Lowland and Olympic Mountains of Washington and the Cascade Mountains of Washington and Oregon. In: Šibrava, V., Bowen, D.Q., Richmond, G. (Eds.), Quaternary Glaciation in the Northern Hemisphere. Quaternary Science Reviews 5, 145–159.Google Scholar
Easterbook, D.J. Advance and retreat of Cordilleran ice sheets in Washington, U.S.A. Géographie physique et Quaternaire 46, (1992). 51–68.Google Scholar
Easterbrook, D.J. Comment on the paper “Determination of 36Cl production rates from the well-dated deglaciation surfaces of Whidbey and Fidalgo Islands, Washington” by T.W. Swanson and M.C. Caffee. Quaternary Research 59, (2003). 132–134.Google Scholar
Easterbrook, D.J., and Kovanen, D.J. Comments on “Marine and limnic radiocarbon reservoir corrections for studies of late- and postglacial environments in Georgia Basin and Puget Lowland, British Columbia, Canada and Washington, USA” by Hutchinson et al. Quaternary Research 63, (2005). 224–226.CrossRefGoogle Scholar
Elisseeff, P., Schmidt, H., Johnson, M., Herold, D., Chapman, N.R., and McDonald, M.M. Acoustic tomography of a coastal front in Haro Strait, British Columbia. Journal of the Acoustical Society of America 106, (1999). 169–184.Google Scholar
Farina, J.M., Salazar, S., Wallem, K.P., Whitman, J.D., and Ellis, J.C. Nutrient exchanges between marine and terrestrial ecosystems: the case of the Galapagos sea lion Zalophus wollebaecki . Journal of Animal Ecology 72, (2003). 873–887.Google Scholar
Fedje, D.W., White, J.M., Wilson, M.C., Nelson, D.E., Vogel, J.S., and Southon, J.R. Vermilion Lakes Site: adaptations and environments in the Canadian Rockies during the latest Pleistocene and early Holocene. American Antiquity 60, (1995). 81–108.Google Scholar
Frison, G.C. Prehistoric Hunters of the High Plains. Second ed. (1991). Academic Press, San Diego, CA. 532 ppGoogle Scholar
Fryxell, J.M., Greener, J., and Sinclair, A.R.E. Why are migratory ungulates so abundant?. American Naturalist 131, (1988). 781–798.CrossRefGoogle Scholar
Fujikawa, W.Y., (2002). A 7600-year Vegetation and Fire History of Mt. Constitution, Orcas Island, Washington, U.S.A. M.S. Thesis, University of Washington, Seattle, WA.Google Scholar
Fuller, W.A., Bayrock, L.A., (1968). Late Pleistocene mammals from central Alberta, Canada. In: Stelck, C.R. (Ed.), Vertebrate Paleontology in Alberta, Report of a Conference, Edmonton, Alberta. University of Alberta, Department of Geology, Bulletin. no. 2, pp. 53–63.Google Scholar
Gates, C.C., and Larter, N.C. Growth and dispersal of an erupting large herbivore population in northern Canada: the Mackenzie wood bison (Bison bison athabascae). Arctic 43, (1990). 231–238.Google Scholar
Geist, V. The relation of social evolution and dispersal in ungulates during the Pleistocene, with emphasis on the Old World deer and the genus Bison . Quaternary Research 1, (1971). 283–315.Google Scholar
Geist, V. Periglacial ecology, large mammals, and their significance to human biology. Bonnichsen, R.A., and Turnmire, K.L. Ice Age People of North America: Environments, Origins, and Adaptations. Center for the Study of the First Americans and Oregon State. (1999). University Press, Corvallis, OR. 75–94.Google Scholar
Gilbow, D.W., (1981). Inference of Human Activity from Faunal Remains. M.A. Thesis (Anthropology), Washington State University, Pullman, WA., 104 pp.Google Scholar
Gordillo, S., and Aitken, A.E. Paleoenvironmental interpretation of late Quaternary marine molluscan assemblages, Canadian Arctic Archipelago. Géographie physique et Quaternaire 54, (2000). 301–315.Google Scholar
Guilbault, J.-P., Barrie, J.V., Conway, K.W., Lapointe, M., and Radi, T. Paleoenvironments associated with the deglaciation process in the Strait of Georgia off British Columbia: microfaunal and microfloral evidence. Quaternary Science Reviews 22, (2003). 839–857.Google Scholar
Gupta, S., Collier, J.S., Palmer-Felgate, A., and Potter, G. Catastrophic flooding origin of shelf valley systems in the English Channel. Nature 448, (2007). 342–345.Google Scholar
Gustafson, C.E., Gilbow, D., and Daugherty, R.D. The Manis Mastodon site: early man on the Olympic Peninsula. Canadian Journal of Archaeology 3, (1979). 157–164.Google Scholar
Guthrie, R.D. Bison evolution and zoogeography in North America during the Pleistocene. Quarterly Review of Biology 45, (1970). 1–15.Google Scholar
Hallett, D.J., Hills, L.V., and Clague, J.J. New accelerator mass spectrometry radiocarbon ages for the Mazama tephra layer from Kootenay National Park, British Columbia. Canadian Journal of Earth Sciences 34, (1997). 1202–1209.Google Scholar
Hansen, H.P. A pollen study of two bogs on Orcas Island, of the San Juan Islands, Washington. Bulletin of the Torrey Botanical Club 70, (1943). 236–243.Google Scholar
Hansen, H.P. Postglacial forest succession, climate, and chronology in the Pacific Northwest. Transactions of the American Philosophical Society 37, (1947). 1–130.CrossRefGoogle Scholar
Harington, C.R. Pleistocene muskoxen (Symbos) from Alberta and British Columbia. Canadian Journal of Earth Sciences, 12, (1975). 903–919.Google Scholar
Harington, C.R., and Morlan, R.E. Evidence for human modification of a Late Pleistocene bison (Bison sp.) bone from the Klondike District, Yukon Territory, Canada. Arctic 55, (2002). 143–147.Google Scholar
Harington, C.R., Ross, R.L.M., Mathewes, R.W., Stewart, K.M., and Beattie, O. A late Pleistocene Steller sea lion (Eumetopias jubatus) from Courtenay, British Columbia: its death, associated biota, and paleoenvironment. Canadian Journal of Earth Sciences 41, (2004). 1285–1297.CrossRefGoogle Scholar
Heaton, T.H., and Grady, F. The Late Wisconsin vertebrate history of Prince of Wales Island, southeast Alaska. Schubert, B.W., Mead, J.I., and Graham, R.W. Ice Age Cave Faunas of North America. (2003). Indiana University Press, Bloomington, IN. 17–53.Google Scholar
Hebda, R.J. British Columbia vegetation and climate history with focus on 6 ka bp. Géographie physique et Quaternaire 49, (1995). 55–79.Google Scholar
Hebda, R.J., and Spalding, D.A.E. Fossils and museums: windows into ancient worlds. Ludvigsen, R. Life in Stone: A Natural History of British Columbia's Fossils. (1996). University of British Columbia Press, Vancouver, BC. 14–24.Google Scholar
Hetherington, R., and Reid, R.G.B. Malacological insights into the marine ecology and changing climate of the late Pleistocene – early Holocene Queen Charlotte Islands archipelago, western Canada, and implications for early peoples. Canadian Journal of Zoology 81, (2003). 626–661.Google Scholar
Hetherington, R., Barrie, J.V., Reid, R.G.B., MacLeod, R., and Smith, D.J. Paleogeography, glacially-induced crustal displacement, and Late Quaternary coastlines on the continental shelf of British Columbia, Canada. Quaternary Science Reviews 23, (2004). 295–318.Google Scholar
Heusser, C.J. Environmental sequence following the Fraser Advance of the Juan de Fuca Lobe, Washington. Quaternary Research 3, (1973). 284–306.Google Scholar
Heusser, C.J., Heusser, L.E., and Peteet, D.M. Late-Quaternary climatic change on the American North Pacific Coast. Nature 315, (1985). 485–487.Google Scholar
Hewitt, A.T., and Mosher, D.C. Postglacial Sea Level History of the Eastern Strait of Juan de Fuca. (1999). American Geophysical Union, Annual Fall Meeting, San Francisco, CA. Abstracts, F586 Google Scholar
Hewitt, A.T., and Mosher, D.C. Late Quaternary stratigraphy and seafloor geology of the eastern Juan de Fuca Strait, British Columbia and Washington. Marine Geology 177, (2001). 295–316.Google Scholar
Hopkins, M.A., Tucker, W., and Mulherrin, N. Simulating ice conditions in Cook Inlet, Alaska. Frederking, R., Kubat, I., and Timco, G. Proceedings of the 16th International Conference on Port and Ocean Engineering under Arctic Conditions, vol. 3. (2001). Canadian Hydraulics Centre, Ottawa, ON. 1261–1264.Google Scholar
Huckleberry, G., Lenz, B., Galm, J., and Gough, S. Recent geoarchaeological discoveries in central Washington. Swanson, T.W. Western Cordillera and Adjacent Areas. (2003). Geological Society of America Field Guide 4, Boulder, CO. 237–249.Google Scholar
Huntley, D.H., Bobrowsky, P.T., and Clague, J.J. Ocean Drilling Program Leg 168S: surficial geology, stratigraphy and geomorphology of the Saanich Inlet area, southeastern Vancouver Island, British Columbia. Marine Geology 174, (2001). 27–41.Google Scholar
Hutchinson, I., James, T.S., Clague, J.J., Barrie, J.V., and Conway, K.W. Reconstruction of late Quaternary sea-level change in southwestern British Columbia from sediments in isolation basins. Boreas 33, (2004). 183–194.Google Scholar
Hutchinson, I., James, T.S., Reimer, P., Bornhold, B., and Clague, J.J. Marine and limnic radiocarbon reservoir corrections for studies of late- and post-glacial environments in the Georgia Basin and Puget Lowland. Quaternary Research 61, (2004). 193–203.Google Scholar
Hutchinson, I., James, T.S., Reimer, P., Bornhold, B., and Clague, J.J. Reply to letter to the editor from Easterbrook and Kovanen. Quaternary Research 63, (2005). 226–227.Google Scholar
Jacobi, R.M., Higham, T.F.G., and Ramsey, C.B. AMS radiocarbon dating of Middle and Upper Paleolithic bone in the British Isles: improved reliability using ultrafiltration. Journal of Quaternary Science 21, (2006). 557–573.Google Scholar
James, T.S., Clague, J.J., Wang, K., and Hutchinson, I. Postglacial rebound at the northern Cascadia subduction zone. Quaternary Science Reviews 19, (2000). 1527–1541.Google Scholar
James, T.S., Hutchinson, I., and Clague, J.J. Improved Relative Sea-level Histories for Victoria and Vancouver from Isolation Basin Coring. (2002). Geological Survey of Canada, Current Research 2002-A16, 7 pp.Google Scholar
Jefferson, G.F., and Goldin, J. Seasonal migration of Bison antiquus from Rancho La Brea. Quaternary Research 31, (1989). 107–112.Google Scholar
Johnsen, T.F., and Brennand, T.A. Late-glacial lakes in the Thompson Basin, British Columbia: paleogeography and evolution. Canadian Journal of Earth Sciences 41, (2004). 1367–1383.Google Scholar
Johnson, S.Y., Dadisman, S.V., Mosher, D.C., Blakely, R.J., Childs, J.R., (2002). Active Tectonics of the Devils Mountain Fault and Related Structures, Northern Puget Lowland and Eastern Strait of Juan de Fuca Region, Pacific Northwest. U.S. Geological Survey Professional Paper 1643. Available online, http://earthquake.usgs.gov/regional/pacnw/activefaults/dmf/index.html#top.Google Scholar
Josenhans, H., Fedje, D.W., Conway, K., and Barrie, J.V. Post-glacial sea levels on the western Canadian continental shelf: evidence for rapid change, extensive subaerial exposure, and early human habitation. Marine Geology 125, (1995). 73–94.Google Scholar
Josenhans, H., Fedje, D.W., Pienitz, R., and Southon, J.R. Early humans and rapidly changing Holocene sea levels in the Queen Charlotte Islands-Hecate Strait, British Columbia, Canada. Science 277, (1997). 71–74.Google Scholar
Keddie, G. The late ice age of southern Vancouver Island. Midden 11, 4 (1979). 16–22.Google Scholar
Kenady, S.M., (1990). Bison letter report to Robert G. Whitlam. On File, Washington Department of Archaeology and Historic Preservation, Olympia.Google Scholar
Kenady, S.M. Evidence of pre-Clovis butchering on a Bison antiquus from the Ayer Pond Site, 45SJ454, in northwest Washington State. (2008). Society for American Archaeology, 73rd Annual Meeting, Vancouver, BC. Canada, 304 Google Scholar
Kenady, S.M., Wilson, M.C., and Schalk, R.F. Indications of butchering on a Bison antiquus from the maritime Pacific Northwest. Current Research in the Pleistocene 24, (2007). 167–170.Google Scholar
Keuler, R.F., (1988). Map Showing Coastal Erosion, Sediment Supply, and Longshore Transport in the Port Townsend 30- by 60-minute Quadrangle, Puget Sound Region, Washington. U.S. Geological Survey Miscellaneous Investigations Series, Map I-1198-E. Scale 1:100,000.Google Scholar
Kokko, H., and López-Sepulcre, A. From individual dispersal to species ranges: perspectives for a changing world. Science 313, (2006). 789–791.Google Scholar
Kovanen, D.J., and Easterbrook, D.J. Extent and timing of Allerød and Younger Dryas age (ca. 12.5–10.0 14C kyr BP) oscillations of the Cordilleran Ice Sheet in the Fraser Lowland, Western North America.. Quaternary Research 57, (2002). 208–224.Google Scholar
Kovanen, D.J., and Easterbrook, D.J. Rooted stumps in the Deming Sand confirmation of abrupt late Pleistocene relative sea level changes in NW Washington. Geological Society of America Abstracts with Programs 32, (2002). 49 Google Scholar
Kovanen, D.J., and Easterbrook, D.J. Paleodeviations of radiocarbon marine reservoir values for the northeast Pacific. Geology 30, (2002). 243–246.Google Scholar
Kurtén, B., Anderson, E., (1980). Pleistocene mammals of North America. Columbia University Press, New York., 443 pp.Google Scholar
Lacourse, T., and Mathewes, R.W. Terrestrial paleoecology of Haida Gwaii and the continental shelf. Fedje, D.W., and Mathewes, R.W. Haida Gwaii: Human History and Environment from the Time of Loon to the Time of Iron People. (2005). University of British Columbia Press, Vancouver, BC. 38–58.Google Scholar
Lacourse, T., Mathewes, R.W., and Fedje, D.W. Paleoecology of late-glacial terrestrial deposits with in situ conifers from the submerged continental shelf of western Canada. Quaternary Research 60, (2003). 180–188.Google Scholar
Larter, N.C., Nishi, J.S., Ellsworth, T., Johnson, D., More, G., and Allaire, D.G. Observations of wood bison swimming across the Liard River, Northwest Territories, Canada. Arctic 56, (2003). 408–412.Google Scholar
Linden, R.H., and Schurer, P.J. Sediment characteristics and sea-level history of Royal Roads anchorage, Victoria, British Columbia. Canadian Journal of Earth Sciences 25, (1988). 1800–1810.Google Scholar
Luternauer, J.L., Conway, K.W., Clague, J.J., and Blaise, B. Late Quaternary geology and geochronology of the central continental shelf of western Canada. Marine Geology 89, (1989). 57–68.Google Scholar
Lyman, R.L. Significance for wildlife management of the late Quaternary biogeography of mountain goats (Oreamnos americanus) in the Pacific Northwest, USA. Arctic and Alpine Research 20, (1988). 13–23.Google Scholar
Marcus, L.F., and Berger, R. The significance of radiocarbon dates for Rancho La Brea. Martin, P.S., and Klein, R.G. Pleistocene Extinctions: A Prehistoric Revolution. (1984). University of Arizona Press, Tucson, AZ. 159–183.Google Scholar
Margalef, R. Perspectives in Ecological Theory. (1968). University of Chicago Press, Chicago, IL.Google Scholar
Matheus, P.E. Diet and co-ecology of Pleistocene short-faced bears and brown bears in Eastern Beringia. Quaternary Research 44, (1995). 447–453.Google Scholar
Matheus, P.E., (2003). Locomotor adaptations and ecomorphology of short-faced Bears (Arctodus simus) in Eastern Beringia. Paleontology Program, Government of the Yukon, Occasional Papers in Earth Sciences no. 7. 126 pp.Google Scholar
Mathewes, R.W., (1985). Paleobotanical evidence for climatic change in southern British Columbia during late-glacial and Holocene time. In: Harington, C.R. (Ed.), Climatic Change in Canada 5: Critical Periods in the Quaternary Climatic History of Northern North America, . Syllogeus 55, pp. 397–422.Google Scholar
Mathewes, R.W. Evidence for Younger Dryas-age cooling on the North Pacific coast of America. Quaternary Science Reviews 12, (1993). 321–331.Google Scholar
Mathews, W.H., Fyles, J.G., and Nasmith, H.W. Postglacial crustal movements in southwestern British Columbia and adjacent Washington State. Canadian Journal of Earth Sciences 7, (1970). 690–702.CrossRefGoogle Scholar
McDonald, H.G. The sloth, the president, and the airport. Washington Geology 26, (1998). 40–42.Google Scholar
McDonald, H.G., Harington, C.R., and De Iuliis, G. The ground sloth Megalonyx from Pleistocene deposits of the Old Crow Basin, Yukon, Canada. Arctic 53, (2000). 213–220.Google Scholar
McDonald, J.N. North American Bison: Their Classification and Evolution. (1981). University of California Press, Berkeley, CA. 316 ppGoogle Scholar
McKenna, M.C. Sweepstakes, filters, corridors, Noah's Arks, and beached Viking funeral ships in paleogeography. Tarling, D.H., and Runcorn, S.K. Implications of Continental Drift to the Earth Sciences. (1973). Academic Press, New York. 295–308.Google Scholar
McNeely, R., Dyke, A.S., Southon, J.R., (2006). Canadian marine reservoir ages, preliminary data assessment. Geological Survey of Canada Open File 5049, 3 pp. + tables.Google Scholar
Meatte, D., (2006). Clovis on the western landscape. 59th Annual Northwest Anthropological Conference, Seattle, WA, Program and Abstracts, 53. Supporting documents on file, Washington Department of Archaeology and Historic Preservation, Olympia, WA.Google Scholar
Meltzer, D.J., and Dunnell, R.C. Fluted points from the Pacific Northwest. Current Research in the Pleistocene 4, (1987). 64–67.Google Scholar
Mosher, D.C., and Hewitt, A.T. Late Quaternary deglaciation and sea-level history of eastern Juan de Fuca Strait, Cascadia. Quaternary International 121, (2004). 23–29.Google Scholar
Myers, J.A., Vellend, M., Gardescu, S., and Marks, P.L. Seed dispersal by white-tailed deer: implications for long-distance dispersal, invasion, and migration of plants in eastern North America. Oecologia 139, (2004). 35–44.Google Scholar
Nagorsen, D.W., and Keddie, G. Late Pleistocene mountain goats (Oreamnos americanus) from Vancouver Island: biogeographic implications. Journal of Mammalogy 81, (2000). 666–675.Google Scholar
Nagorsen, D.W., Keddie, G., and Hebda, R.J. Early Holocene black bears, Ursus americanus, from Vancouver Island. Canadian Field-Naturalist 109, (1995). 11–18.Google Scholar
Overstreet, D.F., Stafford, T.W. Jr. Additions to a revised chronology for cultural and non-cultural mammoth and mastodon fossils in the southwestern Lake Michigan basin. Current Research in the Pleistocene 14, (1997). 70–71.Google Scholar
Peck, T.R., (2004). Bison Ethology and Native Settlement Patterns during the Old Women's Phase on the Northwestern Plains. British Archaeological Reports International Series 1278. 147 pp.Google Scholar
Peterson, K.L., Mehringer, P.J. Jr., and Gustafson, C.E. Late-glacial vegetation and climate at the Manis Mastodon site, Olympic Peninsula, Washington. Quaternary Research 20, (1983). 215–231.Google Scholar
Pigati, J.S., Quade, J., Shahanan, T.M., Haynes, C.V. Jr. Radiocarbon dating of minute gastropods and new constraints on the timing of late Quaternary spring-discharge deposits in southern Arizona, USA. Palaeogeography, Palaeoclimatology, Palaeoecology 204, (2003). 33–45.Google Scholar
Pope, P.H. A fossil bison skull from Prescott, Washington. Northwest Science 26, (1952). 51–54.Google Scholar
Porter, S.C., and Swanson, T.W. Radiocarbon age constraints on rates of advance and retreat of the Puget lobe of the Cordilleran ice sheet during the last glaciation. Quaternary Research 50, (1998). 205–213.Google Scholar
Reimer, P.J., Reimer, R.W., (2001). A marine reservoir correction database and on-line interface. Radiocarbon 43, 11–145. and 461–463.Google Scholar
Rensberger, J.M., Barnosky, A.D., (1978). Letter report to John Cadden, 11 September 1978, from Burke Museum Paleontology Division. Copies on file, Washington Department of Archaeology and Historic Preservation, Olympia.; and Paleontology Division, Burke Museum of Natural History and Culture, Seattle, WA.Google Scholar
Rigg, G.B., and Richardson, C.T. The development of Sphagnum bogs in the San Juan Islands. American Journal of Botany 21, (1934). 610–622.Google Scholar
Ripple, W.J., and Beschta, R.L. Restoring Yellowstone's aspen with wolves. Biological Conservation 138, (2007). 514–519.Google Scholar
Rivals, F., Solounias, N., and Mihlbachler, M.C. Evidence for geographic variation in the diets of late Pleistocene and early Holocene Bison in North America, and differences from the diets of recent Bison . Quaternary Research 68, (2007). 338–346.Google Scholar
Robinson, S.W., and Thompson, G. Radiocarbon corrections for marine shell dates with application to southern Pacific Northwest Coast prehistory. Syesis 14, (1981). 45–57.Google Scholar
Roe, F.G. The North American Buffalo: A Critical Study of the Species in its Wild State. Second ed. (1971). University of Toronto Press, Toronto, ON. 991 ppGoogle Scholar
Schalk, R.F., Kenady, S.M., and Wilson, M.C. Early post-glacial ungulates on the Northwest Coast: implications for hunter-gatherer niches. Current Research in the Pleistocene 24, (2007). 182–185.Google Scholar
Schoeninger, M.J., and DeNiro, M.J. Nitrogen and carbon isotopic composition of bone collagen from marine and terrestrial animals. Geochimica et Cosmochimica Acta 48, (1984). 625–639.Google Scholar
Shackleton, N.J., Duplessy, J.C., Arnold, M., Maurice, P., Hall, M.A., and Cartlidge, J. Radiocarbon age of last glacial Pacific deep water. Nature 335, (1988). 708–711.Google Scholar
Shapiro, B., Drummond, A., Rambaut, A., Wilson, M.C., Sher, A., Pybus, O., Gilbert, M.T.P., Barnes, I., Binladen, J., Willerslev, E., Hansen, A., Baryshnikov, G.F., Burns, J., Davydov, S., Driver, J., Gubin, S.V., Harington, C.R., Keddie, G., Kosintsev, P., Kunz, M.L., Martin, L.D., Stephenson, R., Storer, J., Tedford, R., Vorobiev, A., Zimov, S., and Cooper, A. Rise and fall of the Beringian steppe bison. Science 306, (2004). 1561–1565. and supporting online data Google Scholar
Shaw, J.D., (1972). Late Pleistocene Paleontology of Orcas, Shaw, Lopez, and San Juan Islands of the San Juan Archipelago. M.S. Thesis, University of Washington, Seattle, WA. 60 pp.Google Scholar
Skinner, M.F., and Kaisen, O.C. The fossil Bison of Alaska and preliminary revision of the genus. Bulletin of the American Museum of Natural History 89, (1947). 123–256.Google Scholar
Southon, J., and Fedje, D. A post-glacial record of 14C reservoir ages for the British Columbia coast. Canadian Journal of Archaeology 27, (2003). 95–111.Google Scholar
Sternberg, C.H. Elephas columbi and other mammals in the swamps of Whitman County, Washington. Science n.s. 17, (1903). 511–512.Google Scholar
Stringer, C., (2007). Reaching the extremities: what the early occupation of Britain can tell us about human dispersals. XVII INQUA Congress 2007, Cairns, Australia., Abstract 1484.Google Scholar
Stuiver, M., Pearson, G.W., and Braziunas, T. Radiocarbon age calibration of marine samples back to 9000 cal yr BP. Radiocarbon 28, (1986). 980–1021.Google Scholar
Stuiver, M., Reimer, P.J., Bard, E., Beck, J.W., Burr, G.S., Hughen, K.A., Kromer, B., McCormac, G., Van der Plicht, J., and Spurk, M. INTCAL98 radiocarbon age calibration, 24000–0 cal BP. Radiocarbon 40, (1998). 1041–1084.Google Scholar
Stuiver, M., Reimer, P.J., Reimer, R., (2006). CALIB Radiocarbon Calibration, 5.0.1. Available online, http://calib.qub.ac.uk/calib.Google Scholar
Stutte, N.A., (2004). The Holocene history of bison in the Intermountain West: a synthesis of archaeological and paleontological records from eastern Oregon. M.A. Thesis (Anthropology), Portland State University, Portland, OR., 124 pp.Google Scholar
Swan, P.L., Alexander, S.M., Paquet, P.C., Darimont, C.T., (2003). Biodiversity, focal species and island biogeography on the central coast of British Columbia, Canada: a GIS approach. Fifth International Symposium on GIS and Computer Cartography for Coastal Zone Management, 16–18 October 2003, Genova, Italy., Proceedings. 8 pp.Google Scholar
Swanson, T.W., (2005). Reply to comment by Easterbrook, D.J. (Quaternary Research 2003, 59 #1, 132–134) on “Determination of 36Cl Production Rates from the Well-Dated Deglaciation Surfaces of Whidbey and Fidalgo Islands, Washington.�.� Quaternary Research 63, 228–230.Google Scholar
Swanson, T.W., and Caffee, M.L. Determination of 36Cl production rates from the well-dated deglaciation surfaces of Whidbey and Fidalgo Islands, Washington. Quaternary Research 56, (2001). 366–382.Google Scholar
Talling, P.J., Wynn, R.B., Masson, D.G., Frenz, M., Cronin, B.T., Schiebel, R., Akhmetzhanov, A.M., Dallmeier-Tiessen, S., Benetti, S., Weaver, P.P.E., Georgiopoulou, A., Zühlsdorff, C., and Amy, L.A. Onset of submarine debris flow deposition far from original giant landslide. Nature 450, (2007). 541–544.Google Scholar
Teller, J.T., Murty, T., Nirupama, N., and Chittibabu, P. A possible tsunami in the Labrador Sea related to the drainage of Glacial Lake Agassiz ∼ 8400 years ago. Science of Tsunami Hazards 23, n. 3, (2005). 3–16.Google Scholar
Terborgh, J., Feeley, K., Silman, M., Nuñez, P., and Balukjian, B. Vegetation dynamics of predator-free land-bridge islands. Journal of Ecology 94, (2006). 253–263.Google Scholar
Thompson, T.G., Lorah, J.R., and Rigg, G.B. The acidity of the waters of some Puget Sound bogs. Journal of the American Chemical Society 49, (1927). 2981–2988.Google Scholar
Thomson, R.E., (1981). Oceanography of the British Columbia Coast. Canadian Special Publication of Fisheries and Aquatic Sciences 56, Ottawa., 291 pp.Google Scholar
Thorson, R.M. Ice-sheet glaciation of the Puget Lowland, Washington, during the Vashon stade (Late Pleistocene). Quaternary Research 13, (1980). 303–321.Google Scholar
Thorson, R.M. Glacio-isostatic response of the Puget Sound area, Washington. Geological Society of America Bulletin 101, (1989). 1163–1174.Google Scholar
Thorson, R.M. Earthquake recurrence and glacial loading in western Washington. Geological Society of America Bulletin 108, (1996). 1182–1191.Google Scholar
Tieszen, L.L., and Fagre, T. Effect of diet quality and composition on the isotopic composition of respiratory CO2, bone collagen, bioapatite, and soft tissues. Lambert, J.B., and Grupe, G. Prehistoric Human Bone: Archaeology at the Molecular Level. (1993). Springer-Verlag, New York. 121–155.Google Scholar
Tomlinson, M.S. Quality of waters of San Juan County. Russell, R.H. Geology and Water Resources of the San Juan Islands. (1975). San Juan County Washington, . Washington. Department of Ecology, Office of Technical Services, Water Supply Bulletin 46, Olympia, WA., pp. 127–132.Google Scholar
U.S. EPA, (2002). Methods for Evaluating Wetland Condition: Wetlands Classification. Office of Water, U.S. Environmental Protection Agency, Washington, DC., EPA-822-R-02-107, 36 pp.Google Scholar
Wagner, F.J.E., (1959). Palaeoecology of the Marine Pleistocene Faunas of Southwestern British Columbia. Geological Survey of Canada Bulletin 52. 67 pp.Google Scholar
Waters, M.R., Stafford, T.W. Jr. Redefining the age of Clovis: implications for the peopling of the Americas. Science 315, (2007). 1122–1126.Google Scholar
Webb, S.D., Milanich, J.T., Alexon, R., and Dunbar, J.S. A Bison antiquus kill site, Wacissa River, Jefferson County, Florida. American Antiquity 49, (1984). 384–392.Google Scholar
Whitlock, C. Vegetational and climatic history of the Pacific Northwest during the last 20,000 years: implications for understanding present-day biodiversity. Northwest Environmental Journal 8, (1992). 5–28.Google Scholar
Wilson, M.C. The Casper local fauna and its fossil bison. Frison, G.C. The Casper Site: A Hell Gap Bison Kill on the High Plains. (1974). Academic Press, New York. 124–171.Google Scholar
Wilson, M.C., (1980a). Population dynamics of the Garnsey Site bison. In: Speth, J.D., Parry, W.J. (Eds.), Late Prehistoric Bison Procurement in Southeastern New Mexico: The 1978 Season at the Garnsey Site. Museum of Anthropology, University of Michigan, Technical Report no. 12, Ann Arbor, MI., pp. 88–129.Google Scholar
Wilson, M.C. Morphological dating of late Quaternary bison on the northern plains. Canadian Journal of Anthropology 1, (1980). 81–85.Google Scholar
Wilson, M.C. Late Quaternary vertebrates and the opening of the ice-free corridor, with special reference to the genus Bison . Quaternary International 32, (1996). 97–105.Google Scholar
Wilson, M.C., and Churcher, C.S. The Late Pleistocene Bighill Creek formation and its equivalents in Alberta: correlative potential and vertebrate palaeofauna. Mahaney, W.C. Correlation of Quaternary Chronologies. (1984). GeoBooks, Norwich, UK. 159–175.Google Scholar
Wilson, M.C., Hebda, R.J., Keddie, G., (2003). Early postglacial fossil bison from Vancouver Island, British Columbia, and Orcas Island, Washington: morphology, taxonomy and paleoecological setting. Geological Association of Canada/Mineralogical Association of Canada Joint Annual Meeting, Vancouver, BC., Abstracts, v. 28, 761.Google Scholar
Wilson, M.C., Hills, L.V., Shapiro, B., (2005). Bison “occidentalis” is dead, and the Ice-free Corridor was closed: biogeography, mtDNA, and implications for Paleoindian migrations. 63rd Annual Plains Anthropological Conference, Edmonton, Alberta, Canada, October 19–23, 2005, Programme and Abstracts, 67.Google Scholar
Yorath, C., Kung, R., Franklin, R., (2002). Geoscape Victoria. Geological Survey of Canada, Miscellaneous Report no. 74. Ottawa. Available online, http://geoscape.nrcan.gc.ca/victoria/index_e.php.Google Scholar
Zdanowicz, C.M., Zielinski, G.A., and Germani, M.S. Mount Mazama eruption: calendrical age verified and atmospheric impact assessed. Geology 27, (1999). 621–624.Google Scholar