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A New Index Measuring Evenness

Published online by Cambridge University Press:  11 May 2009

C. Heip
Affiliation:
Department of Zoology, State University of Ghent, Belgium

Extract

The use of diversity and evenness indices is well established in recent ecological literature. Many indices have been proposed, to such an extent that the choice of a suitable index became somewhat of a problem. Recently, however, Hill (1973) introduced a unifying notation where diversity numbers are denned in relation to Rényi's definition of a generalized entropy. Hill showed that his diversity numbers Na of the oth, 1st and 2nd order coincide with three important diversity measures which have been frequently used, No = S, N1 = eH and N2 = 1/SI, where S is the number of species, H is the shannon-Wiener information function –Σpi In pi and SI is Simpson's index . According to this notation, evenness should be calculated by dividing two of Hill's diversity numbers, e.g. N2/N1 = eH/S. This index was proposed by Sheldon (1969), but its use in ecological literature has been negligible. The most commonly used evenness index has been the one proposed by Pielou (1966), e = H/Hmax, with Hmax = lnS. Contrary to Hill's (1973) statement, it shares with Hill's continuum of evenness measures the property of remaining constant when the number of individuals of all species is multiplied with a constant factor. Intuitively, this seems to be a necessary property of an evenness index.

Type
Research Article
Copyright
Copyright © Marine Biological Association of the United Kingdom 1974

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References

REFERENCES

Heip, C. & Engels, P., 1974. Comparing species diversity and evenness indices. Journal of the Marine Biological Association, 54, 559–63.CrossRefGoogle Scholar
Hill, M. O., 1973. Diversity and evenness: a unifying notation and its consequences. Ecology, 54, 427–32.CrossRefGoogle Scholar
Pielou, E. C., 1966. The measurement of diversity in different types of biological collections. Journal of Theoretical Biology, 13, 131–44.CrossRefGoogle Scholar
Sheldon, A. L., 1969. Equitability indices: dependence on the species count. Ecology, 50, 466–7. 36–2CrossRefGoogle Scholar