Hostname: page-component-cd9895bd7-jkksz Total loading time: 0 Render date: 2024-12-28T03:31:40.609Z Has data issue: false hasContentIssue false

Upper Wenlock and lower Ludlow (Silurian), post-extinction graptolites, Všeradice section, Barrandian area, Czech Republic

Published online by Cambridge University Press:  20 May 2016

A. Kozłowska-Dawidziuk
Affiliation:
1Institute of Paleobiology, Polish Academy of Sciences, Twarda 51/55, 00–818 Warszawa, Poland
A.C. Lenz
Affiliation:
2Department of Earth Sciences, University of Western Ontario, London, Ontario, Canada N6A 5B7
P. Štorch
Affiliation:
3Geological Institute AS CR, Rozvojová 135, Praha 6, Suchdol 165 00, Czech Republic

Abstract

The Všeradice section was trenched and continuously sampled for graptolites from the base of the post-extinction upper Homerian strata to low in the Ludlow. The upper Homerian is divisible into three biozones, a lower parvus–nassa Biozone, a middle praedeubeli–deubeli Biozone, and an upper ludensis–gerhardi Biozone, the last named being succeeded by graptolites of the lowest Ludlow nilssoni Biozone. The graptolite diversity of nine species in the upper part of the parvus–nassa Biozone is the highest in the world. Five monograptid species, Pristiograptus parvus Ulst, 1974, P. dubius (Suess, 1851), Colonograptus deubeli (Jaeger, 1959), Colonograptus gerhardi (Kühne, 1955), and Colonograptus praedeubeli (Jaeger, 1990) are described from the upper Homerian. Twelve species of retiolitids, Gothograptus nassa (Holm, 1890), Neogothograptus cf. balticus (Eisenack, 1951), Spinograptus spinosus (Wood, 1900), Spinograptus clathrospinosus (Eisenack, 1951), Spinograptus munchi (Eisenack, 1951), Spinograptus reticulolawsoni (Kozłowska-Dawidziuk, 1997) Spinograptus? cf. nevadensis (Berry and Murphy, 1975), Spinograptus? sp. A and Spinograptus? sp. B, Plectograptus macilentus (Törnquist, 1887), and Plectograptus? karlsteinensis new species and Plectograptus? ovatus new species, are described and illustrated from upper Homerian and lowest Ludlow strata. The two new species occur in the upper parvus–nassa and lowest praedeubeli–deubeli, and ludensis–gerhardi biozones, respectively. Two morphs of Gothograptus nassa, a narrow (more typical) form, and a wide form, are recognized. While overlapping in their overall ranges, their occurrences in any particular small interval are sometimes mutually exclusive, suggesting ecological control.

Type
Research Article
Copyright
Copyright © The Paleontological Society

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Barrande, J. 1850. Graptolites de Bohěme. Prague, 74 p.Google Scholar
Berry, W. B. N and Murphy, M. A. 1975. Silurian and Devonian graptolites of central Nevada. University of California Publications in Geological Sciences, 110:1109.Google Scholar
Bouček, B. 1936. La fauna graptolitique du Ludlowien inférieure de la Bohěme. Rozpravy II (Tčídy české Akademie), 46:137152.Google Scholar
Bouček, B., and Münch, A. 1952. Retioliti stredoevropského svrchního wenlocku a ludlowu. Sborník ústředního ústavu Geologického, XIX. oddíl paleontolický:1151.Google Scholar
Eisenack, A. 1951. Retioliten aus dem Graptolithengestein. Palaeontographica, 100:129163.Google Scholar
Elles, G. L., and Wood, E. M. R. 1908. A Monograph of British graptolites, 62, 273–358. Palaeontographical Society, London,Google Scholar
Elles, G. L., and Wood, E. M. R. 1911. A Monograph of British graptolites, 64, 359–414. Palaeontographical Society, London,Google Scholar
Frech, F. 1897. Lethaea geognostica. Teil 1, Lethaea palaeozoica, I, Graptolitheden. Schweizerbart Stuttgart, 544684.Google Scholar
Gutiérrez-Marco, J. C., Lenz, A. C., Robardet, M., and Piçarra, J. M. 1996. Wenlock-Ludlow graptolite biostratigraphy and extinction: a reassessment from the southwestern Iberian Peninsula (Spain and Portugal). Canadian Journal of Earth Sciences, 33:656663.CrossRefGoogle Scholar
Holland, C. H., Rickards, R. B., and Warren, P. T. 1969. The Wenlock graptolites of the Ludlow district, Shropshire, and their stratigraphical significance. Palaeontology, 12:663683.Google Scholar
Holm, G. 1890. Gotlands graptoliter. Svenska vetenskap-akademie Handlingar, 16:134.Google Scholar
Jaeger, H. 1959. Graptolithen und Stratigraphie des jüngsten Thüringer Silurs. Abhandlugen der deutschen Akademie der Wissenschaften zu Berlin. Klasse fur Chemie, Geologie und Biologie, 2, 197 p.Google Scholar
Jaeger, H. 1990. New geological and biostratigraphical data on the Silurian of SE-Sardinia, p. 101117. In Barca, S., and Jaeger, H., Close affinity with Thuringia. Bolletino della Societa Geologica Italiana, 108.Google Scholar
Jaeger, H. 1991. Neue Standard-Graptolithenzonenfolge nach der “Grossen Krise” an der Wenlock/Ludlow-Grenze (Silur). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 182:303354.CrossRefGoogle Scholar
Jaeger, H., and Robardet, M. 1979. Le Silurien et le Dévonien basal dans le nord de la Province de Seville (Espagne). Geobios, 5:687714.CrossRefGoogle Scholar
Jaekel, O., 1889. Ueber das Alter des sogenanten Graptolithen-Gesteins. Zeitschrift der deutsches geologischen Gesellshaft, 41, 653716.Google Scholar
Koren', T. N. 1991. The lundgreni extinction event in central Asia and its bearing on graptolite biochronology within the Homerian. Proceedings of the Estonian Academy of Sciences, Geology, 40:7478.Google Scholar
Koren', T. N. 1994. The Homerian monograptid fauna of central Asia: zonation, morphology and phylogeny, p. 140148. In Chen, X., Erdtmann, B.-D., and Ni, Y.-N. (eds.), Graptolite Research Today. University Press, Nanjing.Google Scholar
Koren', T. N., and Suyarkova, A. A. 1994. Monograptus deubeli and praedeubeli (Wenlock, Silurian) in the Asian part of the former Soviet Union. Alcheringa, 18:85101.CrossRefGoogle Scholar
Koren', T. N., and Urbanek, A. 1994. Adaptive radiation of monograptids after the Late Wenlock crisis. Acta Palaeontologica Polonica, 39:137167.Google Scholar
Kozłowska-Dawidziuk, A. 1990. The genus Gothograptus (Graptolithina) from the Wenlock of Poland. Acta Palaeontologica Polonica, 35:191209.Google Scholar
Kozłowska-Dawidziuk, A. 1991. Agastograptus from the Mulde Beds of Gotland. Acta Palaeontologica Polonica, 36:143149.Google Scholar
Kozłowska-Dawidziuk, A. 1995. Silurian retiolitids of the East European Platform. Acta Palaeontologica Polonica, 40:261326.Google Scholar
Kozłowska-Dawidziuk, A. 1996. Monograptid and retiolitid graptolites of the Wenlock/Ludlow (Silurian) transition in the East European Platform: biostratigraphic and evolutionary implications. The James Hall Symposium: Second International Symposium on the Silurian System. Program and Abstracts. University of Rochester, N.Y., p. 67.Google Scholar
Kozłowska-Dawidziuk, A. 1997. Retiolitid graptolite Spinograptus from Poland and its membrane structures. Acta Palaeontologica Polonica, 42:391412.Google Scholar
Kozłowska-Dawidziuk, A. 1999. Wydarzenie Cyrtograptus lundgreni a rozwój retiolitów (Graptolithina) na platformie wschodnioeuropejskiej. Przegld Geologiczny, 47:354358.Google Scholar
Kozłowska-Dawidziuk, A., Lenz, A. C., and Storch, P. 1998. Late Homerian-early Ludlow post-extinction graptolites, VŠeradice, Czech Republic, p. 97100. In Gutiérrez-Marco, J. C. and Rábano, I. (eds.), Proceedings 6th International Graptolite Conference (GWG-IPA) & 1998 Field Meeting, IUGS Subcommission on Silurian Stratigraphy Temas Geolégico-Mineros ITGE, Madrid.Google Scholar
KŘíŽ, J. 1992. Silurian field excursions. Prague Basin (Barrandian), Bohemia. National Museum of Wales, Geological Series, 13, Cardiff, 111 p.Google Scholar
KŘíŽ, J., Dufka, P., Jaeger, H., and Schönlaub, H. P. 1993. The Wenlock/Ludlow Boundary in the Prague Basin (Bohemia). Jahrbuch der Geologischen Bundesanstalt, Wien, 136:809839.Google Scholar
Kühne, W. G. 1955. Unterludlow-Graptolithen aus berliner Geschieben. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 100:350401.Google Scholar
Lapworth, C. 1873. On an improved classification of the Rhabdophora . Geological Magazine, 10:500504, 555560.CrossRefGoogle Scholar
Lenz, A. C. 1978. Llandoverian and Wenlockian Cyrtograptus, and some other Wenlockian graptolites from northern and Arctic Canada. Geobios, 11:623653.CrossRefGoogle Scholar
Lenz, A. C. 1993a. Late Wenlock and Ludlow (Silurian) Plectograptinae (retiolitid graptolites), Cape Phillips Formation, Arctic Canada. Bulletins of American Paleontology, 104:152.Google Scholar
Lenz, A. C. 1993b. Late Wenlock-Ludlow (Silurian) extinction, evolution, and biostratigraphy: perspectives from Arctic Canada. Canadian Journal of Earth Sciences, 30:491498.CrossRefGoogle Scholar
Lenz, A. C. 1994a. Extinction and opportunistic evolution among late Wenlock graptolites. Lethaia, 27:111117.CrossRefGoogle Scholar
Lenz, A. C. 1994b. The graptolites “Pristiograptus” praedeubeli (Jaeger) and “Pristiograptus” ludensis (Murchison) (uppermost Wenlock, Silurian) from Arctic Canada: taxonomy and evolution. Canadian Journal of Earth Sciences, 31:14191426.CrossRefGoogle Scholar
Lenz, A. C. 1995. Upper Homerian (Wenlock, Silurian) graptolites and graptolite biostratigraphy, Arctic Archipelago, Canada. Canadian Journal of Earth Sciences, 32:13781392.CrossRefGoogle Scholar
Lenz, A. C., Chen, X., and Y.-N, Ni. 1996. Wenlock, Ludlow and Pridoli? graptolites from Yulin, Guangxi, China. Canadian Journal of Earth Sciences, 33:13901401.CrossRefGoogle Scholar
Moberg, J. C., and Törnquist, S. V. 1909. Retiolitoidea fran Skånes Colonusskrifter. Sveriges Geologiska Undersökning, Sen C, 213:120.Google Scholar
Münch, A. 1942. Die Graptolithenfauna des unteren Ludlows. Beiträge zur geologie von Thüringens. Jena.Google Scholar
Murchison, R. I. 1839. The Silurian System. London, xxxii + 768 p.Google Scholar
Ni, Y.-N. 1997. Late Homerian (Wenlock, Silurian) graptolites from Shidian, western Yunnan, China. Acta Palaeontologica Sinica, 36:310320 (Chinese with English summary).Google Scholar
Obut, A. M., and Zaslavskaya, N. M. 1979. Razvitie rabdosom i ekologiya Gothograptus nassa (Holm), p. 3033. In Sreda i Zhihn v Geologicheskom Proshlom. Nauka, Novosibirsk.Google Scholar
Obut, A. M., and Zaslavskaya, N. M. 1983. Semyestva retiolitid i ikh filogeneticheskie otnosheniya, p. 103113. In Dagys, A. S., and Koren, T. N.’ (eds.), Morfologiya sistematika bespozvonochnykh fanerozoa. Akademia Nauk SSSR, Sibirskoe otdelenie. Institut Geologie i Geofiziki, Izdatel'stvo “Nauk”, Moscow,Google Scholar
Paškevičius, J. 1979. Biostratigrafiya i graptolity Silura Litvy. Vilnius Mokslas Publishers, 268 p.Google Scholar
Patočka, F. Vlasimsky, P., and Blechova, K. 1993. Geochemistry of Early Paleozoic volcanics of the Barrandian Basin (Bohemian Massif, Czechoslovakia): implications for paleotectonic reconstructions. Jahrbuch der Geologischen Bundesanstalt, Wien, 136:873896.Google Scholar
PŘíbyl, A. 1942. Revision der Pristiograpten aus den Untergattugen Colonograptus n. subg. and Saetograptus n. subg. Mitteilungen der tschechischen Akademie der Wissenschaften, 52:122.Google Scholar
PŘíbyl, A. 1943. Revision aller Vertreter der Gattung Pristiograptus aus der Gruppe P. dubius und P. vulgaris aus dem böhmischen und ausländischen Silur. Mitteilungen der tschechischen Akademie der Wisseschaften, Jahrgang. 53:149.Google Scholar
Rickards, R. B. 1965. New Silurian graptolites from the Howgill Fells (Northern England). Palaeontology, 8:247271.Google Scholar
Rickards, R. B. 1995. Utility and precision of Silurian graptolite biozones. Lethaia, 28:129137.CrossRefGoogle Scholar
Rickards, R. B., and Wright, A. J. 1997. Graptolite zonation in the late Wenlock (Early Silurian), with a new graptolite-brachiopod fauna from New South Wales. Records of the Australian Museum, 49:229248.CrossRefGoogle Scholar
Rickards, R. B., Packham, G. H., Wright, A. J., and Williamson, P. L. 1995. Wenlock and Ludlow graptolite faunas and biostratigraphy of the Quarry Creek district, New South Wales. Association of Australasian Palaeontologists Memoir 17, 68 p.Google Scholar
Salvador, A. (ed.). 1994. International Stratigraphic Guide, (second edition). International Union of Geological Sciences and Geological Society of America.Google Scholar
Scotese, C. R., and Golonka, J. 1992. PALEOMAP Paleogeographic Atlas. PALEOMAP Progress Report #20, Department of Geology, University of Texas at Arlington.Google Scholar
Štorch, P. 1994. Graptolite biostratigraphy of the Lower Silurian (Llandovery and Wenlock) of Bohemia. Geological Journal, 29:137165.CrossRefGoogle Scholar
Suess, E. 1851. Über böhmische Graptoliten. Naturwissenschaftliche Abhandlungen von W. Haidinger, 4:87134.Google Scholar
Tomczyk, H. 1956. Wenlock i ludlow w synklinie kieleckiej Gór Swietokrzyskich. Prace Instytutu Geologicznego, 16:177.Google Scholar
Tomczyk, H. 1974. Profil glebokich otworow wiertniczych Instytutu Geologicznego, Bartoszyce IG 1, Goldap IG 1. Wydawnictwa Geologiczne, 14:122137.Google Scholar
Törnquist, S. L. 1887. Anteckningar om de äldre Paleozoiska Leden i Ostthuringen och Vogtland. Geologiska Föreningens Forhandlingar, 9:471492.CrossRefGoogle Scholar
Ulst, R. ZH. 1974. Posledovatel'nost’ pristiograptov v pogranichnykh otlozheniyakh venlocka i ludlova v sredniy Pribaltike, p. 105122. In Obut, A. M. (ed.), Graptolites of the USSR. Publishing House “Nauka”. Siberian Branch, Novosibirsk.Google Scholar
Urbanek, A. 1958. Monograptidae from erratic boulders of Poland. Palaeontologia Polonica, 9:1105.Google Scholar
Waterlot, G. 1945. Les graptolites du Maroc. Service géologique du Maroc, Notes et Mémoires, 63:45103.Google Scholar
Wiman, C. 1895. Über die Graptoliten. Bulletin of the Geological Institution of the University of Uppsala, 2:239316.Google Scholar
Wood, E. M. R. 1900. The Lower Ludlow formation and its graptolite fauna. Quarterly Journal of the Geological Society of London, 56:415492.CrossRefGoogle Scholar