Hostname: page-component-cd9895bd7-jkksz Total loading time: 0 Render date: 2024-12-27T11:03:39.487Z Has data issue: false hasContentIssue false

Observations on the Early Cambrian helicoplacoid echinoderms

Published online by Cambridge University Press:  20 May 2016

J. W. Durham*
Affiliation:
Department of Integrative Biology and Museum of Paleontology, University of California, Berkeley 94720

Abstract

The Early Cambrian helicoplacoid echinoderms occur in the Cordilleran Geosyncline of western North America in strata correlated with the Atdabanian Stage of Siberia. Several higher taxa are recognized on the basis of inferred differences in the water vascular system, test organization, and external morphology. These are subclass Polyplacida Durham, with genus Polyplacus Durham; subclass Helicoplacida Durham and Caster, with n. family Helicoplacidae, type genus Helicoplacus Durham and Caster (with tubefeet emerging between two contemporaneous ambulacral plates); n. family Westgardellidae, with type n. genus Westgardella, type species H. curtisi (Durham and Caster) (with tubefeet emerging between two sequential ambulacral plates). The genus Waucobella Durham is also referred to Westgardellidae. Helicoplacus gilberti Durham and Caster, H. everndeni Durham, H. casteri n. sp., H. guthi n. sp., H. sp. a, and H. sp. b are assigned to Helicoplacidae. The genus Westgardella includes H. firbyi Durham, 1967, and W. blancoensis n. sp., in addition to the type species. No evidence of flooring plates separating the radial water vessel from the interior of the test is recognized. The mouth is at the top of the test in the interpretation adopted herein and not lateral as inferred by others; therefore, the ambulacral system is not triradiate. Illustration identified as Helicoplacus curtisi by Paul and Smith includes misidentified plates and should not be referred to this species.

Type
Research Article
Copyright
Copyright © The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Broadhead, T. W., and Waters, J. A. 1980. Echinoderms. Notes for a short course. University of Tennessee, Department of Geological Sciences, Studies in Geology, 3:1235.Google Scholar
Derstler, K. 1981. Morphological diversity of Early Cambrian echinoderms, p. 7175. In Taylor, M. E. (ed.), Short Papers for the Second International Symposium on the Cambrian System. U.S. Geological Survey, Open File Report 81-743, 254 p. Google Scholar
Durham, J. W. 1964a. The Helicoplacoida and some possible implications. Yale Scientific Magazine, 39:2425, 28.Google Scholar
Durham, J. W. 1964b. Echinodermata, p. 207208. 1964 Yearbook of Science and Technology. McGraw-Hill, New York.Google Scholar
Durham, J. W. 1967. Notes on the Helicoplacoidea and early echinoderms. Journal of Paleontology, 41:97102.Google Scholar
Durham, J. W. 1987. On Helicoplacus (Helicoplacoida, Echinodermata). Sixth International Echinoderm Conference, Victoria, British Columbia, Canada, August 23–28, 1987. Scientific Program, Abstracts [pages not numbered].Google Scholar
Durham, J. W. 1988. On Helicoplacus (Helicoplacoida, Echinodermata), p. 794. In Burke, R. D., Miladenov, P. W., Lambert, P., Parsley, P., and R. L. (eds.), Echinoderm Biology. Proceedings of the Sixth International Echinoderm Conference, Victoria, 23–28 August 1987. A. A. Balkema, Rotterdam.Google Scholar
Durham, J. W., and Caster, K. E. 1963. Helicoplacoidea: a new class of echinoderms. Science, 140:820822.Google Scholar
Durham, J. W., and Caster, K. E. 1966. Helicoplacoids, p. U131136. In Moore, R. C. (ed.), Treatise on Invertebrate Paleontology, Pt. U, Echinodermata 3(1). Geological Society of America and University of Kansas Press, Lawrence.Google Scholar
Fritz, W. H. 1972. Lower Cambrian trilobites from the Sekwi Formation type section, Mackenzie Mountains, northwestern Canada. Bulletin of the Geological Survey of Canada, 212:158.Google Scholar
Lawrence, J. M. 1987. A Functional Biology of Echinoderms. The Johns Hopkins University Press, Baltimore, 340 p.Google Scholar
McMenamin, M. A. S. 1987. The emergence of animals. Scientific American, 255:94102.Google Scholar
Moore, J. N. 1976. Depositional environments of lower Paleozoic rocks in the White-Inyo Range, Inyo County, California, p. 112. In Moore, J. N. and Fritsche, A. E. (eds.), Depositional Environments of Lower Paleozoic Rocks in the White-Inyo Mountains, Inyo County, California. Pacific Coast Paleogeography Field Guide 1, Pacific Section of the Society of Economic Paleontologists and Mineralogists, Los Angeles.Google Scholar
Nelson, C. A. 1962. Lower Cambrian–Precambrian succession, White-Inyo Mountains, California. Geological Society of America Bulletin, 73:139144.Google Scholar
Nelson, C. A. 1966. Geologic map of the Blanco Mountain quadrangle, Inyo and Mono Counties, California. U.S. Geological Survey, Geologic Quadrangle Map 6Q–529.Google Scholar
Nelson, C. A. 1976. Late Precambrian–Early Cambrian stratigraphic and faunal succession of Eastern California and the Precambrian–Cambrian boundary, p. 3142. In Moore, J. N. and Fritsche, A. E. (eds.), Depositional Environments of Lower Paleozoic Rocks in the White-Inyo Mountains, Inyo County, California. Pacific Coast Paleogeography Field Guide 1, Pacific Section of the Society of Economic Paleontologists and Mineralogists, Los Angeles.Google Scholar
Nelson, C. A., and Durham, J. W. 1966. Guidebook for field trip to Precambrian–Cambrian succession, White-Inyo Mountains, California, November 17–20, 1966. Privately printed, Los Angeles, 15 p.Google Scholar
Nichols, D. 1966. Functional morphology of the water-vascular system, p. 219244. In Boolootian, R. A. (ed.), Physiology of Echinodermata. Interscience Publishers, New York.Google Scholar
Paul, C. R. C. 1977. Evolution of primitive echinoderms, p. 123158. In Hallam, A., (ed.), Patterns of Evolution as Illustrated by the Fossil Record. Elsevier, Amsterdam.Google Scholar
Paul, C. R. C., and Smith, A. B. 1984. The early radiation and phylogeny of echinoderms. Biological Reviews of the Cambridge Philosophical Society, 59:443481.Google Scholar
Smith, A. B. 1984. Classification of the Echinodermata. Palaeontology, 27:431459.Google Scholar
Smith, A. B. 1988a. Fossil evidence for the relationships of extant echinoderm classes and their times of divergence, p. 8597. In Paul, C. R. C. and Smith, A. B. (eds.), Echinoderm Phylogeny and Evolutionary Biology. Oxford University Press, New York.Google Scholar
Smith, A. B. 1988b. Patterns of diversifications and extinction in early Palaeozoic echinoderms. Palaeontology, 31:799828.Google Scholar
Sprinkle, J. 1973. Morphology and evolution of blastozoan echinoderms. Museum Comparative Zoology, Harvard University, Special Publication, 284 p.Google Scholar
Sprinkle, J. 1976a. Classification and phylogeny of “pelmatozoan” echinoderms. Systematic Zoology, 25:8391.Google Scholar
Sprinkle, J. 1976b. Biostratigraphy and paleoecology of Cambrian echinoderms from the Rocky Mountains. Brigham Young University Geological Studies, 23:6173.Google Scholar
Sprinkle, J. 1981. Diversity and evolutionary patterns of Cambrian echinoderms, p. 219221. In Taylor, M. E. (ed.), Short Papers for the Second International Symposium on the Cambrian System. U.S. Geological Survey Open File Report.Google Scholar
Sprinkle, J. 1983. Patterns and problems in echinoderm evolution, p. 118. In Jangoux, M. and Lawrence, J. M. (eds.), Echinoderm Studies, 1, A. A. Balkema, Rotterdam.Google Scholar
Wainwright, S. A., Biggs, W. D., Currey, J. D., and Gosline, J. M. 1982. Mechanical Design in Organisms. Princeton University Press, Princeton, New Jersey, 423 p.Google Scholar