Published online by Cambridge University Press: 20 May 2016
Until the late 1960's, most of Nicholson's types of Paleozoic bryozoans were not available for study. We present a set of coded characters of many of Nicholson's types, which should assist in bringing his species into conformity with current taxonomic standards so that his species can be consistently recognized and used in biostratigraphic, paleobiogeographic, and evolutionary studies.
Cladistic and phenetic analyses of these species permit comparisons between inferred phylogenies of Nicholson specimens, adaptive morphospace, and treatise-based systematic relationships. Specimen-based cladistic and phenetic analyses of Nicholson's species both produce clusters that are congruent with existing family-level taxonomic assignments of species in the collection. However, cladistic analysis more fully retrieves the pattern of branching, or degree of relatedness, among higher taxa. Phenetic clusters represent adaptive peaks in morphospace for these specimens, but higher level “phenons” are strongly affected by multiple evolution of the same character states.