Hostname: page-component-78c5997874-t5tsf Total loading time: 0 Render date: 2024-11-13T00:48:20.292Z Has data issue: false hasContentIssue false

Aulacopleurid trilobites from the Upper Ordovician of Virginia

Published online by Cambridge University Press:  20 May 2016

Jonathan M. Adrain*
Affiliation:
Department of Geoscience, University of Iowa, 121 Trowbridge Hall, Iowa City 52242,

Abstract

Five species of aulacopleurid trilobites occur in rich, silicified trilobite faunas from the Upper Ordovician of Virginia: Harpidella triloba (Hu, 1975a), Strasburgaspis cona (Hu, 1971), and Strasburgaspis? n. sp. A, all from the Turinian Edinburg Formation, Harpidella whittingtoni new species, from the overlying Turinian Oranda Formation, and Harpidella evitti new species, from the Chatfieldian Martinsburg Formation. The species of Harpidella, similar to other sets of congeneric taxa known from the formations, are subtly but pervasively differentiated. In addition to qualitative differentia such as the expression of the eye socle and of pygidial tubercle rows, the species are shown to differ in cranidial and librigenal shape via landmark-based geometric morphometric analysis. The genus Strasburgaspis (type species S. cona) is new. It is assigned to Aulacopleuridae on the basis of its micropygous morphology, but it shares potential apomorphies with Brachymetopidae and could prove to be the most plesiomorphic representative of that family.

Type
Research Article
Copyright
Copyright © The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Adrain, J. M. 1996. A new otarionine trilobite from the Henryhouse Formation (Silurian, Ludlow) of Oklahoma. Journal of Paleontology, 70:611614.Google Scholar
Adrain, J. M., and Chatterton, B. D. E.. 1994. The aulacopleurid trilobite Otarion, with new species from the Silurian of northwestern Canada. Journal of Paleontology, 68:305323.Google Scholar
Adrain, J. M., and Chatterton, B. D. E.. 1995a. The otarionine trilobites Harpidella and Maurotarion, with species from northwestern Canada, the United States, and Australia. Journal of Paleontology, 69:307326.Google Scholar
Adrain, J. M., and Chatterton, B. D. E.. 1995b. Aulacopleurine trilobites from the Llandovery of northwestern Canada. Journal of Paleontology, 69:326340.Google Scholar
Adrain, J. M., and Chatterton, B. D. E.. 1996. The otarionine trilobite Cyphaspis, with new species from the Silurian of northwestern Canada. Journal of Paleontology, 70:100110.Google Scholar
Adrain, J. M., and Edgecombe, G. D.. 1996. Devonian aulacopleurid trilobites of the Malvinokaffric Realm. Geobios, 29:417436.Google Scholar
Adrain, J. M., and Fortey, R. A.. 1997. Ordovician trilobites from the Tourmakeady Limestone, western Ireland. Bulletin of the Natural History Museum, London, Geology Series, 53:79115.Google Scholar
Adrain, J. M., and Kloc, G. J.. 1997. Lower Devonian aulacopleuroidean trilobites from Oklahoma. Journal of Paleontology, 71:703712.Google Scholar
Alberti, G. K. B. 1969. Trilobiten des jüngeren Siluriums sowie des unter- und Mitteldevons. I. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 520:1692.Google Scholar
Angelin, N. P. 1854. Palaeontologica Scandinavica. Pars II, Crustacea formationis transitionis. Academiae Regiae Scientarum Suecanae (Holmiae), 2592.Google Scholar
Barrande, J. 1846. Notice préliminaire sur le systême Silurien et les Trilobites de Bohême. Hirschfeld, Leipzig, 97 p.Google Scholar
Burmeister, H. 1843. Die Organisation der Trilobiten aus ihren lebenden Verwandten entwickelt; nebst einer systematischen Uebersicht aller zeither beschriebenen Arten. Reimer, Berlin, 147 p.Google Scholar
Campbell, K. S. W. 1977. Trilobites of the Haragan, Bois d'Arc and Frisco Formations (Early Devonian), Arbuckle Mountains Region, Oklahoma. Oklahoma Geological Survey Bulletin, 123:1139.Google Scholar
Chatterton, B. D. E. 1971. Taxonomy and ontogeny of Siluro-Devonian trilobites from near Yass, New South Wales. Palaeontographica Abteilung A, 137:1108.Google Scholar
Chatterton, B. D. E. 1994. Ordovician proetide trilobite Dimeropyge, with a new species from northwestern Canada. Journal of Paleontology, 68:541556.Google Scholar
Chatterton, B. D. E., Siveter, D. J., Edgecombe, G. D., and Hunt, A. S.. 1990. Larvae and relationships of the Calymenina (Trilobita). Journal of Paleontology, 64:255277.Google Scholar
Chatterton, B. D. E., Edgecombe, G. D., Speyer, S. E., Hunt, A. S., and Fortey, R. A.. 1994. Ontogeny and relationships of Trinucleoidea (Trilobita). Journal of Paleontology, 68:523540.Google Scholar
Cooper, B. N. 1953. Trilobites from the Lower Champlainian Formations of the Appalachian Valley. Geological Society of America Memoir, 55, 69 p.Google Scholar
Cooper, B. N., and Cooper, G. A.. 1946. Lower Middle Ordovician stratigraphy of the Shenandoah Valley, Virginia. Geological Society of America Bulletin, 57:35114.Google Scholar
Cooper, G. A. 1956. Chazyan and related brachiopods. Smithsonian Miscellaneous Collections, 127:11245.Google Scholar
Dean, W. T. 1966. The lower Ordovician stratigraphy and trilobites of the Landreyan valley and the neighbouring district of the Montagne Noire, southwestern France. Bulletin of the British Museum (Natural History), Geology Series, 12:247353.Google Scholar
Evitt, W. R. 1951. Some Middle Ordovician trilobites of the families Cheiruridae, Harpidae and Lichidae. Journal of Paleontology, 25:587616.Google Scholar
Evitt, W. R. 1961. Early ontogeny in the trilobite family Asaphidae. Journal of Paleontology, 35:986995.Google Scholar
Evitt, W. R., and Tripp, R. P.. 1977. Silicified Middle Ordovician trilobites from the families Encrinuridae and Staurocephalidae. Palaeontographica Abteilung A, 157:109174.Google Scholar
Fortey, R. A., 1997. Late Ordovician trilobites from southern Thailand. Palaeontology, 40:397449.Google Scholar
Fortey, R. A., and Owens, R. M.. 1975. Proetida—a new order of trilobites. Fossils and Strata, 4:227239.Google Scholar
Fortey, R. A., Harper, D. A. T., Ingham, J. K., Owen, A. W., Parkes, M. A., Rushton, A. W. A., and Woodcock, N. H.. 2000. A revised correlation of Ordovician rocks in the British Isles. The Geological Society, Special Report, 24, 83 p.Google Scholar
Hall, J. 1847. Palaeontology of New York, Volume 1, Containing descriptions of the organic remains of the lower division of the New York System (equivalent to the Lower Silurian rocks of Europe). Van Benthuysen, Albany, 338 p.Google Scholar
Hammann, W. 1992. The Ordovician trilobites from the Iberian Chains in the province of Aragón, NE-Spain. I. The trilobites of the Cystoid Limestone (Ashgill Series). Beringeria, 6:3219.Google Scholar
Hu, C.-H. 1971. Ontogeny and sexual dimorphism of lower Paleozoic Trilobita. Palaeontographica Americana, 7:31155.Google Scholar
Hu, C.-H. 1974a. Ontogenies of a few Middle Ordovician trilobites from the Edinburg Formation of Virginia, U.S.A. Bulletin of the Geological Survey of Taiwan, 24:5167.Google Scholar
Hu, C.-H. 1974b. Ontogenies of two Middle Ordovician trilobites from the Edinburg Formation, Virginia. Transactions and Proceedings of the Palaeontological Society of Japan, 95:353363.Google Scholar
Hu, C.-H. 1975a. Ontogenies of four species of silicified Middle Ordovician trilobites from Virginia. Proceedings of the Geological Society of China, 18:115127.Google Scholar
Hu, C.-H. 1975b. Ontogenies of two species of silicified trilobites from Middle Ordovician, Virginia. Transactions and Proceedings of the Palaeontological Society of Japan, n.s., 97:3247.Google Scholar
Hu, C.-H. 1976. Ontogenies of three species of silicified Middle Ordovician trilobites from Virginia. Transactions and Proceedings of the Palaeontological Society of Japan, n.s., 101:247263.Google Scholar
Hughes, N. C., and Chapman, R. E.. 1995. Growth and variation in the Silurian proetide trilobite Aulacopleura konincki and its implications for trilobite palaeobiology. Lethaia, 28:333353.Google Scholar
Leslie, S. A. 2000. Mohawkian (Upper Ordovician) conodonts of eastern North America and Baltoscandia. Journal of Paleontology, 74:11221147.Google Scholar
Leslie, S. A., and Bergström, S. M.. 1995. Timing of the Trenton Transgression and revision of the North American late Middle Ordovician stage classification based on K-bentonite bed correlation, p. 4954. In Cooper, J. D., Droser, M. L., and Finney, S. C. (eds.), Ordovician Odyssey: Short Papers for the Seventh International Symposium on the Ordovician System. Pacific Section Society for Sedimentary Geology (SEPM), Fullerton, California.Google Scholar
Leslie, S. A., and Bergström, S. M.. 1997. Use of K-bentonite beds as time-planes for high resolution lithofacies analysis and assessment of net rock accumulation rate: An example from the upper Middle Ordovician of eastern North America, p. 1121. In Klapper, G., Murphy, M. A., and Talent, J. A. (eds.), Paleozoic Sequence Stratigraphy, Biostratigraphy, and Biogeography: Studies in Honor of J. Granville (“Jess”) Johnson. Geological Society of America Special Paper, 321.Google Scholar
Lespérance, P. J., and Weissenberger, J. A. W.. 1998. Trilobites of the Pabos Formation (Ashgillian, Upper Ordovician), Percé area, Quebec. Journal of Paleontology, 72:303316.Google Scholar
Lütke, F. 1980. Zur Evolution der altpaläozoischen Proetina (Trilobita). Senckenbergiana lethaea, 61:73144.Google Scholar
M'Coy, F. 1846. A Synopsis of the Silurian Fossils of Ireland. Dublin, 72 p.Google Scholar
M'Coy, F. 1849. On the classification of some British fossil Crustacea with notices of some forms in the University collection at Cambridge. Annals and Magazine of Natural History, 2(4):161179, 330–335, 392–414.Google Scholar
Öpik, A. A. 1937. Trilobiten aus Estland. Tartu Ülikooli Geoloogia-Intituudi Toimetused, 52:1163.Google Scholar
Owens, R. M. 1973. British Ordovician and Silurian Proetidae (Trilobita). Monographs of the Palaeontographical Society, 535:198.Google Scholar
Owens, R. M., and Hammann, W.. 1990. Proetide trilobites from the Cystoid Limestone (Ashgill) of NW Spain, and the suprageneric classification of related forms. Paläontologische Zeitschrift, 64:221244.Google Scholar
Owens, R. M., and Thomas, A. T.. 1975. Radnoria, a new Silurian proetacean trilobite, and the origins of the Brachymetopidae. Palaeontology, 18:809822.Google Scholar
Pillet, J. 1969. La classification des Proetidae (Trilobites). Bulletin de la Société d'Études Scientifiques de l'Anjou, n.s., 7:5378.Google Scholar
Prantl, F., and Pribyl, A.. 1951. A revision of the Bohemian representatives of the Family Otarionidae R. & E. Richter (Trilobitae). Sborník Státního Geologického Ústavu Československé Republiky, oddíl paleontologický, 17:353512. (In Czech and English, with Russian summary; for 1950)Google Scholar
Přibyl, A. 1946. Příspěvek k poznání Českých proetidû (Trilobitae). Rozpravy České Akademie Věd a Uměni, 55(10):137.Google Scholar
Přibyl, A., and Vaněk, J.. 1981a. Studie zur Morphologie und Phylogenie der familie Otarionidae R. & E. Richter (Trilobita). Palaeontographica Abteilung A, 173:160208.Google Scholar
Repina, L. N., Yaskovich, B. V., Askarina, N. A., Petrunina, Z. E., Poniklenko, I. A., Rubanov, D. A., Bolgova, G. V., Golikov, A. N., Khairullina, T. I., and Posochova, M. M.. 1975. Stratigrafiya i fauna nizhnego paleozoya sevrn'ch predgorij Turkestanskogo i Alajskogo chrevtov (juzhn'j Tyan'-Shan) [Stratigraphy and fauna of the Lower Paleozoic of the southern submontane belt of Turkestan and the Alai ridges (southern Tien-Shan)]. Trudy Instituta Geologii i Geofiziki. Sibirskoe Otdelenie, Akademiya Nauk SSSR, 278:1351. (In Russian)Google Scholar
Richter, R., and Richter, E.. 1926. Die Trilobiten des Oberdevons. Beiträge zur Kenntnis devonsicher Trilobiten IV. Abhandlungen der Preussischen Geologischen Landesanstalt, n.f., 99:1314.Google Scholar
Sheets, H. D. 2003. IMP Suite. Software distributed by the author at: http://www.canisius.edu/~sheets/morphsoft.html.Google Scholar
Tripp, R. P., and Evitt, W. R.. 1981. Silicified Lichidae (Trilobita) from the Middle Ordovician of Virginia. Geological Magazine, 118:665677.Google Scholar
Tripp, R. P., and Evitt, W. R.. 1983. Silicified trilobites of the genus Dimeropyge from the Middle Ordovician of Virginia, p. 229240. In Briggs, D. E. G. and Lane, P. D. (eds.), Trilobites and other early arthropods. Papers in honour of Professor H. B. Whittington, F. R. S. Special Papers in Palaeontology, 30.Google Scholar
Tripp, R. P., and Evitt, W. R.. 1986. Silicified trilobites of the family Asaphidae from the Middle Ordovician of Virginia. Palaeontology, 29:705724.Google Scholar
Tripp, R. P., Rudkin, D. R., and Evitt, W. R.. 1997. Silicified trilobites of the genus Sphaerocoryphe from the Middle Ordovician of Virginia. Canadian Journal of Earth Sciences, 34:770788.Google Scholar
Vaněk, J., and Vokáč, V.. 1997. Discovery of Harpidella McCoy, 1849 (Trilobita) in the Kralodvorian (Upper Ordovician) near Levín (Prague Basin, Czech Republic). Palaeontologia Bohemiae, 3:79.Google Scholar
Whittington, H. B. 1941. Silicified Trenton trilobites. Journal of Paleontology, 15:492522.Google Scholar
Whittington, H. B. 1956. Silicified Middle Ordovician trilobites: The Odontopleuridae. Bulletin of the Museum of Comparative Zoology, Harvard University, 114:155288.Google Scholar
Whittington, H. B. 1959. Silicified Middle Ordovician trilobites: Remopleurididae, Trinucleidae, Raphiophoridae, Endymionidae. Bulletin of the Museum of Comparative Zoology, Harvard, 121:371496.Google Scholar
Whittington, H. B. 1963. Middle Ordovician trilobites from Lower Head, western Newfoundland. Bulletin of the Museum of Comparative Zoology, Harvard, 129:1118.Google Scholar
Whittington, H. B., and Campbell, K. S. W.. 1967. Silicified Silurian trilobites from Maine. Bulletin of the Museum of Comparative Zoology, Harvard, 135:447483.Google Scholar
Whittington, H. B., and Evitt, W. R.. 1954. Silicified Middle Ordovician trilobites. Geological Society of America Memoir, 59, 59 p.Google Scholar
Zenker, J. C. 1833. Beiträge zur Naturgesichte der Urwelt. Organische Reste (Petrefacten) aus der Altenbruger Braunkohlen-Formation dem Blankenburger Quadersandstein, Jenaischen bunten Sandstein und Böhmischen Uebergangsgebirge. Jena, 67 p.Google Scholar
Zhang, W.-T. 1974. [Silurian Trilobita], p. 173187. In Nanking Institute of Geology and Paleontology, Academica Sinica (ed.), [Handbook of the Stratigraphy and Paleontology of Southwest China]. Science Press, Beijing. (In Chinese; published as Chang, W.-T.)Google Scholar