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Systematic position of the Paulchoffatiinae (Multituberculata, Mammalia)

Published online by Cambridge University Press:  14 July 2015

David W. Krause
Affiliation:
Departments of Anatomical Sciences and Earth and Space Sciences, State University of New York, Stony Brook 11794
Gerhard Hahn
Affiliation:
Institut für Geologie und Paläontologie der Philipps-Universität, Universitäts-Gebiet Lahnberge, Hans-Meerwein-Strasse, D-3550 Marburg, Bundesrepublik Deutschland

Extract

The late Jurassic Paulchoffatiinae, one of two named subfamilies of the family Paulchoffatiidae (suborder Plagiaulacoidea), are generally recognized as the most primitive known, undoubted multituberculates (e.g., Hahn, 1969; Clemens and Kielan-Jaworowska, 1979; Hahn and Hahn, 1983; Hahn et al., 1989). Despite the primitive morphology of members of this subfamily (Paulchoffatia and Pseudobolodon), three features have been posited as autapomorphic for the subfamily (or family), and thus to preclude it from the ancestry of later multituberculates: 1) the distribution of cusps on M2 (Hahn, 1969, 1971); 2) the complex structure of I3 (Hahn, 1969, 1971); and 3) the position of M2 relative to M1 (Van Valen, 1976; Clemens and Kielan-Jaworowska, 1979; Hahn, 1987). Van Valen (1976) argued that the first two features are not unequivocally autapomorphic for paulchoffatiines and that they could, and indeed should, be considered primitive for the order. Hahn (1977) responded to Van Valen's assessment but Clemens and Kielan-Jaworowska (1979, p. 138) opined that “neither argument is particularly compelling” in light of the poor early record of multituberculates and the consequent lack of knowledge concerning morphological variability.

Type
Taxonomic Notes
Copyright
Copyright © The Journal of Paleontology 

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