A knowledge of the exact length of the breeding season for improved breeds under different environmental conditions in this country is lacking. Such knowledge is of importance so that the lambing season can be arranged according to farm conditions. Such knowledge, apart from providing the necessary background for the successful application of artificial insemination and administration of hormones, helps to explain certain problems of infertility. By selection it should be possible to build up strains, by extending the breeding season at one or both ends, so as to get lambing at any required time. As the results below show, crossbreeding and selection could lead to early sexual maturity, prolonged breeding season or restricted lactation anoestrus, which would undoubtedly increase the reproductive potentiality of the ewe.

During the course of three breeding seasons, daily observations of oestrus were undertaken on the following groups of ewes, all of which were running with ochred rams. One hundred and twenty ewes (pure bred and first-cross) of different breeds and ages, running with vasectomized rams under natural conditions. Twenty-eight grade Suffolk ewes fed on a submaintenance diet. Eighteen grade Suffolk ewes running with a fertile ram. Twenty ewes of several breeds exposed to artificial light.

The effects of heredity (breed and individual), environment (season, year and nutrition), age and artificial light on the breeding season and related phenomena have been investigated. The results and conclusions were as follows:

1. (a) There are breed differences in the extent of the breeding season, cycle length, incidence of silent heat and duration of heat (Table 29). (b) The duration of the breeding season is related to the geographical origin (latitude and altitude) of the breed, (c) The duration of the breeding season of the first-cross is intermediate between that of the two parents, (d) Individual differences in the number of oestrous cycles per ewe per season were more marked in the mountain breeds.

2. (a) Of the ewe lambs, 79% exhibited oestrus during the shortest days only of the first breeding season. Their breeding season is not spread evenly about the shortest day as it is with adults. (b) The occurrence and length of the breeding season in ewe lambs is associated with early birth dates or with higher growth rates.

3. (a) Significant differences exist between breeds in the age at first oestrus. (b) Ewe lambs born early in the season showed their first oestrus at later age and heavier weight than those born late.

4. Annual differences in the duration of the breeding season, cycle length, incidence of silent heat and occurrence of first oestrus were negligible.

5. (a) Submaintenance diet had no effect on the onset of the breeding season, but it converted oestrus into silent heat. Conception occurred less frequently after periods of underfeeding. (b) At high latitudes nutrition has only a minor effect on the breeding season of the ewe.

6. (a) A constant high ratio of artificial darkness (8 hr. light: 16 hr. darkness) hastened the onset of the breeding season some 57 and 27 days in the two experimental groups. (b) A constant high ratio of artificial light (16 hr. light: 8 hr. darkness) hastened the end of the breeding season some 104 days on an average (in one experimental group), (c) There were breed differences in the latency of initiation and of cessation of the induced breeding season, (d) In the induced breeding season cycles of ovulation preceded the first oestrus, (e) Two thresholds of pituitary activity are suggested, one for the onset of ovulation, and the other for the manifestation of oestrus.

7. (a) A high frequency of cycles outside the normal range (14–19 days) was observed in the mountain breeds and in ewe lambs. (b) The shortest average cycle length coincided with the shortest days of the year.

8. A high frequency of silent heats (during the breeding season) was observed in the mountain breeds, in ewe lambs and during the second half of the breeding season.

9. Oestrus was of longer duration in adults and yearlings than in ewe lambs.

10. (a) Post-partum heat occurred in 56% of Suffolk ewes with an average lactation anoestrus of 35 days. (b) There is a relationship between the early onset of the breeding season and the incidence of post-partum heat, (c) Conception post-partum is partly inhibited by lactation.

11. Mating behaviour was observed in animals of several breeds and ages (three patterns were recorded for ewes and eight patterns for rams).

12. Irregular columnar cells in the vaginal smear of the ewe characterized late pregnancy.

In addition, the breeding season of wild and domestic sheep was reviewed in relation to the environment with special reference to the length of daylight.

Permanent buildings of a variety of designs have been in widespread use for many years as potato stores on the Continent of Europe and in the United States of America. (For a summary of recent literature, with references, see Rose & Cook, 1949.) There are examples of buildings being used as potato stores in this country (e.g. Keith, 1941), but the bulk of the crop is stored in clamps. Because the climates of the U.S.A. and the Continent of Europe differ markedly from that of Great Britain, it is not possible to use the types of store developed in those regions as models for British stores. Over much North America and the Continent of Europe the stores are designed to withstand severe winter frosts, and the main problem is to keep the temperature of the tubers high enough to prevent freezing, the metabolic heat being readily removed if necessary by a small amount of controlled ventilation. Under British conditions the mean daily temperature for most of the storage season, above the optimum for potato storage (40·42° F.), and therefore the main problem is that of keeping the tubers cool. This could, of course, be achieved by using a cold store, but cold storage is considered too expensive. To design a store, information is required about the temperatures attained by stacks of potatoes at different times of the year, the effectiveness ventilation in controlling the temperature, the relation of temperature in different parts of the stack to dimensions of the stack, and to the insulation both of the bin and of the building. To obtain this data a programme of work was initiated, and the results of the first year's work in a building of very low insulating capacity are described in this paper. The climatic conditions during this year, i.e. 1946–7, were very abnormal, a very cold winter followed by a hot spell in the spring which provided an abnormally severe test of an uninsulated store. Later papers will consider storage in an insulated building with adequate ventilation.

In three groups of field experiments totalling fiftysix observations phosphate responses were found to be very significantly and inversely related both to the percentage saturation of the b.e.c. and exchangeable calcium as a percentage of exchangeable calcium plus hydrogen. The availability of the native soil phosphate was significantly and directly related to these values. The relationships between phosphate response or availability and the amounts of acid-soluble, adsorbed and water-soluble phosphate in the soils were much less conclusive.

In acid soils considerable amounts of phosphate can be retained in a plant-available form associated with exchangeable bases, mainly calcium. Since practically all the soils from the field experiments lay within the range pH 4·7–6·3, it is believed that this form of phosphate is operative in these soils. The presence of such phosphate in the soil is, however, a function of the degree of base saturation in so far as this governs (probably through its influence on soil pH) the equilibrium distribution of the native soil phosphate between the available base-linked (saloid-bound) form and the less available colloid-bound form. With decreasing pH and base saturation increasing amounts of colloid-bound phosphate are formed at the expense of the saloid-bound phosphate. The degree of base saturation is therefore directly related to phosphate availability and, consequently, inversely related to phosphate response.

In view of the significant regressions obtained, and the association of the exchangeable bases with a specific form of available phosphate, it is suggested that until methods are worked out for determining the amount of saloid-bound phosphate exchangeable base data may be used for assessing phosphato availability and response in acid soils. This method is contrasted with the more conventional methods of determining acid-soluble and adsorbed phosphates which are not always specific regarding tho form of phosphate extracted and the values for which, since they may represent the sum of a number of forms of different availabilities, aro difficult to interpret.

It is assumed that maximum growth requires maximum transpiration, and that maximum transpiration can be maintained by keeping the soil near to field capacity throughout the growing season. Transpiration rates can be calculated from weather data (the basic principles are outlined and an example of the calculation given), and the paper describes four field experiments in which attempts were made to control the water content of the soil throughout the growing season, by irrigation from overhead spray-lines.

In spite of differences in season and soil, the four sets of data are consistent in showing that maximum sugar yield is obtained when the soil-moisture deficit (amount of rain or irrigation needed restore the soil to field capacity) does not exceed about 2 in. in mid-July, or about 4 in. in mid-September.

The effects, on induction and maintenance of oestrus and on pelt changes, were compared for various schedules of entirely artificial light of constant intensity.

A period of exposure to low environmental temperature was without effect on subsequent response to light treatment.

In one experiment, after periods on 6 hr. artificial light or on daylight, groups were put upon, daily, (i) 7 hr. light as 6 hr. + 5 hr. dark + 1 hr. light, (ii) the same total amount of light but divided as 6 hr. + 1 hr. separated by darkness increasing by 5 min. each day, and (iii) light increasing from 7 hr. by 5 min. daily. Groups (ii) and (iii) became oestrous together, later than group (i).

Subsequently all were placed on 7 hr. total light daily, applied in various ways. If given as a single continuous period daily, oestrus ended and moult into winter coat occurred. The latent period depended upon the treatment previously imposed. The duration of oestrus on treatment (i) was greater than normal.

In another experiment females were put on, daily, (iv) 24 hr. light, (v) 14 hr. light, and (vi) 4 hr. light as 2 hr. + 10 hr. dark + 2 hr. light. First response was seen on (vi), but on the average the quickest was on (v) and slowest on (iv). Thus darkness contributes to the induction of oestrus.

Possible interpretations are discussed.

1. In continuation of experiments made in 1949 (Holmes et al. 1950) an experiment was carried out from 7 May until 23 September 1950, to compare close-folding and rotational grazing of cows on pastures which were liberally treated with nitrogenous fertilizer. With close-folding the cows were moved daily to an area of fresh pasture calculated to supply the day's feed requirements; the rate of stocking for the day ranged from thirty to sixty-five cows per acre. With rotational grazing the cows were stocked on pasture at the rate of seven to eight cows per acre and moved from one pasture to the other at intervals of 3–4 days. Two uniform groups of six Ayrshire cows were used in a double reversal layout with four periods each of 5 weeks. The same pastures as in 1949—a permanent pasture and a cocksfoot ley—suitably divided by electric fences were used. No supplementary feeding was given.

2. The average yield per acre from close-folding was 241 cow-days, 732 gal. of milk and 320 lb. live-weight gain—equivalent to 632 lb. digestible crude protein and 4316 lb. starch equivalent. Rotational grazing on similar adjoining paddocks gave 181 cow-days, 557 gal. of milk and 285 lb. liveweight gain per acre, equivalent to 486 lb. digestible crude protein and 3371 lb. starch equivalent.

3. The average daily milk yield per cow was 29·5 lb. for one group and 30·2 lb. for the other. For close-folding it was 29·6 lb. and for rotational grazing it was 30·1 lb., none of the differences being significant. Nor were any differences in the average live weight of the groups or in their live-weight gains significant.

4. The increased production per acre from closefolding compared with rotational grazing—amounting to 20–40%—could be related to the increased efficiency with which the available pasture was consumed.

5. Close-folding had no harmful effect on the pastures. Although the season was exceptionally wet, poaching occurred only in the gates and alleyways. 6. The frequent applications of nitrogenous fertilizer throughout the season maintained regular production of good herbage. In June and July, however, despite the fertilizer applied, the crudeprotein content dropped on some paddocks to 13% of the dry matter, a figure barely sufficient to maintain high milk yields.

7. Comparison of the production per acre in 1950 with that in 1949 where close-folding was used, showed an increase of 34% in starch equivalent utilized on the permanent pasture. For each extra cwt. ‘Nitro-Chalk’ applied in 1950 over that in 1949, 190 lb. starch equivalent were produced. On the cocksfoot the response to additional nitrogen was reduced because of potash deficiency induced by cropping for grass-drying in 1948.

8. The factors affecting the increased production from close-folding and the fertilizer requirements of pasture are discussed.

1. The experiments started in 1949 to determine the weight and chemical composition of the herbage dry matter eaten by two groups of dairy cows grazing under a rotational and a close-folding system of management have been continued in 1950.

2. Good-quality herbage was fed in adequate quantity to both groups for 20 weeks, in four fiveweekly periods allowing a double change-over system for the groups. The weights of protein, fibre, other extractives, lignin and minerals eaten daily by the close-folded group were determined. Difficulty in obtaining truly representative herbage samples from the rotational paddock made such detailed analyses for the rotationally grazed cattle not worth while.

3. The weight of dry matter eaten by the closefolded cows was remarkably constant, the mean was 26 lb./cow/day and the standard deviation ±2·7 lb. The weight eaten by the rotationally grazed cows was more difficult to measure and appeared to vary considerably, the average being 31 lb. dry matter/ cow/day, and the standard deviation ±3·1 lb. Reasons are suggested why this figure of 31 lb. may be at least 5% too high.

4. The relationship between the chemical composition of herbage as cut for sampling and the probable composition of the herbage as eaten by the grazing animal is discussed. Correction curves are given to convert the protein and fibre contents of herbage as cut in these experiments to values corresponding to those in the herbage as eaten.

5. An attempt has been made to provide a balance sheet, in terms of D.C.P. and S.E., of animal production against nutrient intake. A deficiency of both protein and energy existed during the first spring period in both 1949 and 1950 (according to presentday standards), and although the protein balance became positive later in the season, the energy provided by the grass eaten was still barely adequate.

6. Mineral balances calculated for the closefolded cows showed deficiencies of both calcium and phosphorus during the period of high milk yield, with the balances becoming positive later in the grazing season.