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The age, fauna and palaeoenvironment of the Late Triassic fissure deposits of Tytherington, South Gloucestershire, UK

Published online by Cambridge University Press:  17 September 2007

D. I. WHITESIDE  and
J. E. A. MARSHALL
D. I. WHITESIDE*
Affiliation:
14 Crown Road, Chelsfield, Orpington, Kent BR6 6JN, UK
J. E. A. MARSHALL
Affiliation:
School of Ocean and Earth Science, National Oceanography Centre Southampton, University of Southampton, Waterfront Campus, European Way, Southampton, SO14 3ZH, UK
*
*Author for correspondence: [email protected]
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Abstract

Important vertebrate faunas occur in fissure deposits of Late Triassic–Jurassic age in SW Britain. Although the faunas are well described, their age and palaeoenvironment remain poorly understood. One such fissure system was documented in detail during quarrying operations at Tytherington and has yielded in situ palynomorphs that add much information concerning its age and palaeoenvironment. Significantly, the Tytherington fauna is of the sauropsid type that has generally been dated as Norian or pre-Penarth Group transgression and was also regarded as representing a distinct upland fauna. The palynomorphs, which include a significant marine component, demonstrate that the Tytherington Triassic fissures are infilled with Late Triassic (Rhaetian) sediments that match specific levels in the Westbury Formation. In addition, many of the Tytherington solutional fissures probably formed during the Rhaetian and are consistent with a fluctuating saline to freshwater environment. There is no prima facie evidence of solutional formation and infilling of the reptile-bearing deposits before the Rhaetian trangression. The fissure reptile fauna, which includes the early dinosaur Thecodontosaurus, inhabited a small fire-swept limestone island in the Rhaetian sea. The features of the herpetofauna are entirely consistent with this island model which has Quaternary analogues.

Keywords

palynomorphsRhaetianreptilestransgressiondinosaursfissuresislands
Type
Original Article
Information
Geological Magazine , Volume 145 , Issue 1 , January 2008 , pp. 105 - 147
Copyright
Copyright © Cambridge University Press 2007

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References

Achilles, H. 1981. Die Rätische und Liassische mikroflora Frankens. Palaeontographica Abteilung B 179, 1–86.Google Scholar
Allison, P. A. & Wright, V. P. 2005. Switching off the carbonate factory: A-tidality, stratification and brackish wedges in epeiric seas. Sedimentary Geology 179, 175–84.CrossRefGoogle Scholar
Arnold, E. N. 1976. Fossil reptiles from Aldabra Atoll, Indian Ocean. Bulletin of the British Museum of Natural History (Zoology) 29, 85–116.Google Scholar
Audley-Charles, M. G. 1970. Triassic palaeogeography of the British Isles. Quarterly Journal of the Geological Society of London 126, 49–89.CrossRefGoogle Scholar
Batten, D. J. & Grenfell, H. R. 1996. Green and blue-green algae. In Palynology: principles and applications (eds Jansonius, J. & McGregor, D. C.), pp. 205–14. American Association of Stratigraphic Palynologists Foundation.Google Scholar
Batten, D. J., Koppelhus, E. B. & Nielsen, L. H. 1994. Uppermost Triassic to Middle Jurassic palynofacies and palynomiscellanea in the Danish Basin and Fennoscandian Border Zone. Cahiers de Micropaléontologie N.S. 9, 21–45.Google Scholar
Below, R. 1987. Evolution und Systematik von Dinoflagellaten-Zysten aus der Ordnung Peridiniales. I. Allgemeine Grundlagen und Subfamilie Rhaetogonyaulacoideae (Familie Peridiniaceae). Palaeontographica Abteilung B 205, 1–164.Google Scholar
Benfield, A. C. & Warrington, G. 1988. New records of the Westbury Formation (Penarth Group, Rhaetian) in North Yorkshire, England. Proceedings of the Yorkshire Geological Society 47, 29–32.Google Scholar
Bennett, A. F. & Gorman, G. C. 1979. Population density and energetics of lizards on a tropical island. Oecologia 42, 339–58.Google Scholar
Benton, M. J. 1993. Reptilia. In The fossil Record 2 (ed. Benton, M. J.), pp. 681–715. London: Chapman & Hall.Google Scholar
Benton, M. J. 1994. Late Triassic to Middle Jurassic extinctions among continental tetrapods: testing the pattern. In In the shadow of the dinosaurs (eds Fraser, N. C. & Sues, H.-D.), pp. 366–97. New York: Cambridge University Press.Google Scholar
Benton, M. J. & Clark, J. M. 1988. Archosaur phylogeny and the relationships of the Crocodylia. In The Phylogeny and Classification of the Tetrapods. Vol. 1: Amphibians, Reptiles and Birds (ed. Benton, M. J.), pp. 295–338. Oxford: Clarendon Press.Google Scholar
Benton, M. J., Juul, L., Storrs, G. W. & Galton, P. M. 2000. Anatomy and systematics of the prosauropod dinosaur Thecodontosaurus antiquus from the Upper Triassic of Southwest England. Journal of Vertebrate Paleontology 20, 77–108.CrossRefGoogle Scholar
Benton, M. J. & Spencer, P. S. 1995. British Triassic fossil reptile sites. In Fossil Reptiles of Great Britain (eds Wimbledon, W. A. & Palmer, D.), pp. 33–95. London: Chapman & Hall.CrossRefGoogle Scholar
Black, M. 1980. Andros island, chalk and oceanic oozes. Unpublished work of Maurice Black (eds Jeans, C. V. & Rawson, P. E.), pp. 1–100. Yorkshire Geological Society Occasional Publication 5.Google Scholar
Bonaparte, J. F. & Sues, H.-D. 2006. A new species of Clevosaurus (Lepidosauria: Rhynchocephalia) from the Upper Triassic of Rio do Sul, Brazil. Palaeontology 49, 917–23.CrossRefGoogle Scholar
Braithwaite, C. J. R., Taylor, J. D. & Kennedy, W. J. 1973. The Evolution of an atoll: the depositional and erosional history of Aldabra. Philosophical Transactions of the Royal Society B 266, 307–40.Google Scholar
Bravo, T. 1953. Lacerta maxima n. sp. de la fauna continental extinguida en el Pleistoceno de las Islas Canarias. Estudios Geológicos Instito de Investigaciones geológicas Lucas Mallado 9, 7–34.Google Scholar
Bristowe, A. & Raath, M. A. 2004. A juvenile coelophysoid skull from the Early Jurassic of Zimbabwe, and the synonymy of Coelopysis and Syntarsus. Palaeontologica Africana 40, 31–41.Google Scholar
Brodkorb, P. 1959. Pleistocene birds from New Providence Island Bahamas. Bulletin of the Florida State Museum 9, 349–71.Google Scholar
Case, T. J. 1975. Species numbers, density compensation, and colonizing ability of lizards on islands in the Gulf of California. Ecology 56, 3–18.CrossRefGoogle Scholar
Chidley, T. R. E. & Lloyd, J. W. 1977. A mathematical model study of fresh water lenses. Ground Water 15, 215–22.Google Scholar
Choquette, P. W. & James, N. P. 1988. Introduction. In Paleokarst (eds James, N. P. & Choquette, P. W.), pp. 1–21. Springer Verlag.Google Scholar
Ciofi, C. 1999. The Komodo Dragon. Scientific American 280, 92–9.CrossRefGoogle Scholar
Colbert, E. H. 1958. Triassic tetrapod extinction at the end of the Triassic Period. Proceedings of the National Academy Science USA 44, 973–7.Google Scholar
Crush, P. J. 1984. A late upper Triassic sphenosuchid crocodilian from Wales. Palaeontology 27, 131–57.Google Scholar
Cuny, G. 1995. French vertebrate faunas and the Triassic–Jurassic boundary. Palaeogeography, Palaeoclimatology, Palaeoecology 119, 343–58.CrossRefGoogle Scholar
Cuny, G. 2004. A Late Triassic cynodont from Holwell Quarries (Somerset, England). Oryctos 5, 69–73.Google Scholar
Curtis, M. T. 1981. The mineralization of Southfields Quarry, Chipping Sodbury, Avon. Proceedings of the Bristol Naturalists’ Society 40, 36–44.Google Scholar
Davis, J. W. 1881. Notes on the Fish remains of the Bone-Bed at Aust near Bristol, with the description of some new genera and species. Quarterly Journal of the Geological Society of London 37, 414–26.CrossRefGoogle Scholar
Donovan, D. T., Curtis, M. T. & Curtis, S. A. 1989. A psiloceratid ammonite from the supposed Triassic Penarth Group of Avon, England. Palaeontology 32, 231–5.Google Scholar
Duffin, C. J. 1980. The Upper Triassic section at Chilcompton, Somerset, with notes on the Rhaetic of the Mendips in general. Mercian Geologist 7, 251–67.Google Scholar
Duffin, C. J. 1995. The first Sphenodontian remains (Lepidosauromorpha, Reptilia) from the Late Triassic of the Gaume (southern Belgium). Bulletin de la Société belge de Géologie 104, 35–41.Google Scholar
Edwards, R. A. 1999. The Minehead district – a concise account of the geology. British Geological Survey. London: The Stationery Office, 128 pp.Google Scholar
Etheridge, R. 1964. Late Pleistocene lizards from Barbuda, British West Indies. Bulletin of the Florida State Museum (Biological Sciences) 9, 4.Google Scholar
Etheridge, R. 1966. Pleistocene lizards from New Providence. Quarterly Journal of the Florida Academy of Science 28, 349–58.Google Scholar
Evans, S. E. 1980. The skull of a new eosuchian reptile from the Lower Jurassic of South Wales. Zoological Journal of the Linnean Society 70, 203–64.CrossRefGoogle Scholar
Evans, S. E. & Kermack, K. A. 1994. Assemblages of small tetrapods from the early Jurassic of Britain. In In the shadow of the dinosaurs (eds Fraser, N. C. & Sues, H.-D.), pp. 271–82. New York: Cambridge University Press.Google Scholar
Evans, S. E., Prasad, G. V. R. & Manhas, B. K. 2001. Rhynchocephalians (Diapsida: Lepidosauria) from the Jurassic Kota Formation of India. Zoological Journal of the Linnean Society 133, 285–301.CrossRefGoogle Scholar
Fisher, J., Simon, N. & Vincent, J. 1969. The Red Book. Wildlife in danger. London: Collins, 368 pp.Google Scholar
Ford, D. 1988. Characteristics of Dissolutional Cave Systems in Carbonate Rocks. In Paleokarst (eds James, N. P. & Choquette, P. W.), pp. 25–57. Springer Verlag.CrossRefGoogle Scholar
Francis, E. H. 1959. The Rhaetic of the Bridgend District, Glamorganshire. Proceedings of the Geologists’ Association 70, 158–70.Google Scholar
Fraser, N. C. 1982. A new rhynchocephalian from the British Upper Triassic. Palaeontology 25, 709–25.Google Scholar
Fraser, N. C. 1985. Vertebrate faunas from Mesozoic fissure deposits of South West Britain. Modern Geology 9, 273–300.Google Scholar
Fraser, N. C. 1986 a. Terrestrial reptiles at the Triassic–Jurassic boundary in South West Britain. Modern Geology 10, 147–57.Google Scholar
Fraser, N. C. 1986 b. New Triassic sphenodontids from South-West England and a review of their classification. Palaeontology 29, 165–86.Google Scholar
Fraser, N. C. 1988 a. The osteology and relationships of Clevosaurus (Reptilia: Sphenodontida). Philosophical Transactions of the Royal Society B 321, 125–78.Google Scholar
Fraser, N. C. 1988 b. Latest Triassic terrestrial vertebrates and their biostratigraphy. Modern Geology 13, 125–40.Google Scholar
Fraser, N. C. 1993. A new sphenodontian from the Early Mesozoic of England and North America: implications for correlating early Mesozoic continental deposits. In The Nonmarine Triassic (eds Lucas, S. G. & Morales, M.), pp. 135–9. New Mexico Museum of Natural History and Science Bulletin 3.Google Scholar
Fraser, N. C. 1994. Assemblages of small tetrapods from British Late Triassic fissure deposits. In In the shadow of the dinosaurs (eds Fraser, N. C. & Sues, H.-D.), pp. 214–26. New York: Cambridge University Press.Google Scholar
Fraser, N. C., Padian, K., Walkden, G. M. & Davis, A. L. M. 2002. Basal dinosauriform remains from Britain and the diagnosis of the Dinosauria. Palaeontology 45, 79–95.CrossRefGoogle Scholar
Fraser, N. C. & Walkden, G. M. 1983. The ecology of a Late Triassic reptile assemblage from Gloucestershire, England. Palaeogeography, Palaeoclimatology, Palaeoecology 42, 341–65.CrossRefGoogle Scholar
Fraser, N. C. & Walkden, G. M. 1984. Two Late Triassic terrestrial ecosystems from South West England. In Third Symposium on Mesozoic Terrestrial Ecosystems, Short Papers (eds Reif, W.-E. & Westphal, F.), pp. 87–92. Tubingen (ATTEMPTO Verlag).Google Scholar
Fraser, N. C., Walkden, G. M. & Stewart, V. 1985. The first pre-Rhaetic therian mammal. Nature 314, 161–3.CrossRefGoogle Scholar
Frost, D. V. 1998. Geology of the country around Northallerton. Memoir for 1:50000 Geological Sheet 42 (England and Wales). London: The Stationery Office, 109 pp.Google Scholar
Gardiner, B. G. 1961. New Rhaetic and Liassic beetles. Palaeontology 4, 87–9.Google Scholar
Godefroit, P., Cuny, G., Delsate, D. & Roche, M. 1998. Late Triassic Vertebrates from Syren (Luxembourg). Neues Jahrbuch für Geologie und Palaontologie Abhandlungen 210, 305–43.CrossRefGoogle Scholar
Halstead, L. B. & Halstead, J. 1981. Dinosaurs. Poole, Dorset, UK: Blandford Press, 170 pp.Google Scholar
Halstead, L. B. & Nicoll, P. G. 1971. Fossilized caves of Mendips. Studies in Speleology 2, 93–102.Google Scholar
Hamilton, D. 1962. Some notes on the Rhaetic sediments of the Filton By-Pass substitute, near Bristol. Proceedings of the Bristol Naturalist Society 30, 279–85.Google Scholar
Harris, T. M. 1957. A Liasso-Rhaetic flora in South Wales. Proceedings of the Royal Society of London B 147, 289–308.Google Scholar
Harris, T. M. 1958. Forest fire in the Mesozoic. Journal of Ecology 46, 447–53.Google Scholar
Harrison, W. J. 1876. On the occurrence of the Rhaetic Beds in Leicestershire. Quarterly Journal of the Geological Society of London 32, 212–18.CrossRefGoogle Scholar
Heckert, A. B., Hunt, A. P. & Lucas, S. G. 1996. Redescription of Redondasuchus reseri, a Late Triassic aetosaur (Reptilia: Archosauria) from New Mexico (USA) and the biochronology and phylogeny of aetosaurs. Geobios 29, 619–32.Google Scholar
Hesselbo, S. P., Robinson, S. A. & Surlyk, F. 2004. Sea-level change and facies development across potential Triassic–Jurassic boundary horizons, SW Britain. Journal of the Geological Society, London 161, 365–79.CrossRefGoogle Scholar
Hounslow, M. W., Posen, P. E. & Warrington, G. 2004. Magnetostratigraphy and biostratigraphy of the Upper Triassic and lowermost Jurassic succession, St. Audrie's Bay, UK. Palaeogeography, Palaeoclimatology, Palaeoecology 213, 331–58.CrossRefGoogle Scholar
Irmis, R. B. 2004. First report of Megapnosaurus (Theropoda: Coelophysoidea) from China. PaleoBios 24, 11–18.Google Scholar
Jakucs, L. 1977. Morphogenesis of Karst regions: variants of Karst evolution. Adam Hilger (Bristol) and Akadémiai Kiadó (Budapest), 284 pp.Google Scholar
Jones, B. 1992 a. Caymanite, a cavity-filling deposit in the Oligocene–Miocene Bluff Formation of the Cayman Islands. Canadian Journal of Earth Science 29, 720–36.CrossRefGoogle Scholar
Jones, B. 1992 b. Void-filling deposits in karst terrains of isolated oceanic islands: a case study from Tertiary carbonates of the Cayman Islands. Sedimentology 39, 857–76.Google Scholar
Jones, T. R. 1862. A monograph of the fossil Estheridae. Monograph of the Palaeontographical Society 14, 1–134.CrossRefGoogle Scholar
Kellaway, G. A. & Welch, F. B. A. 1993. Geology of the Bristol District. London: British Geological Survey, HMSO, 199 pp.Google Scholar
Kermack, D. 1984. New prosauropod material from South Wales. Zoological Journal of the Linnean Society of London 82, 101–17.CrossRefGoogle Scholar
Kermack, K. A., Mussett, F. & Rigney, H. W. 1973. The lower jaw of Morganucodon. Zoological Journal of the Linnean Society 53, 87–175.CrossRefGoogle Scholar
Korth, W. W. 1979. Taphonomy of microvertebrate fossil assemblages. Annals of the Carnegie Museum 48, 235–85.CrossRefGoogle Scholar
Kühne, W. G. 1956. The Liassic Therapsid Oligokyphus. London: British Museum (Natural History), 149 pp.Google Scholar
Long, R. A. & Murry, P. A. 1995. Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4, 254 pp.Google Scholar
Lucas, S. G. 1998. Global Triassic tetrapod biostratigraphy and biochronology. Palaeogeography, Palaeoclimatology, Palaeoecology 143, 347–84.CrossRefGoogle Scholar
Lucas, S. G. 1999. Tetrapod-based correlation of the nonmarine Triassic. Zentralblatt für Geologie und Paläontologie Teil 1 (Heft 7–8), 497–521.Google Scholar
Lucas, S. G., Heckert, A. B., Fraser, N. C. & Hubert, P. 1999. Aetosaurus from the Upper Triassic of Great Britain and its biochronological significance. Neues Jahrbuch für Geologie und Palaontologie Monatshefte 9, 568–76.CrossRefGoogle Scholar
Lucas, S. G., Heckert, A. B. & Huber, P. 1998. Aetosaurus (Archosauromorpha) from the Upper Triassic of the Newark Supergroup, eastern United States, and its biochronological significance. Palaeontology 41, 1215–30.Google Scholar
Lucas, S. G. & Hunt, A. 1994. The chronology and palaeobiogeography of mammalian origins. In In the shadow of the dinosaurs (eds Fraser, N. C. & Sues, H.-D.), pp. 335–51. New York: Cambridge University Press.Google Scholar
Lucas, S. G. & Tanner, L. H. 2004. Late Triassic extinction events. Albertiana 31, 31–40.Google Scholar
Lucas, S. G. & Tanner, L. H. 2006. Tetrapod biostratigraphy and biochronology of the Late Triassic–Jurassic transition on the southern Colorado Plateau, USA. Palaeogeography, Palaeoclimatology, Palaeoecology 244, 242–56.CrossRefGoogle Scholar
Lund, J. J. 1977. Rhaetic to Lower Liassic palynology of the onshore south-eastern North Sea Basin. Danmarks Geologiske Undersøgelse II 109, 1–129.Google Scholar
Luo, Z. & Wu, X.-C. 1994. The small tetrapods of the Lower Lufeng Formation. Yunnan, China. In In the shadow of the dinosaurs (eds Fraser, N. C. & Sues, H.-D.), pp. 251–70. New York: Cambridge University Press.Google Scholar
MacArthur, R. H. & Wilson, E. O. 1963. An equilibrium theory of insular zoogeography. Evolution 17, 373–87.CrossRefGoogle Scholar
Marshall, J. E. A. & Whiteside, D. I. 1980. Marine influence in the Triassic “uplands”. Nature 287, 627–8.CrossRefGoogle Scholar
Martill, D. M. & Dawn, A. 1986. Fossil vertebrates from new exposures of the Westbury Formation (Upper Triassic) at Newark, Nottinghamshire. Mercian Geologist 10, 127–33.Google Scholar
Mayall, M. J. 1979. The Clay mineralogy of the Rhaetic transgression in Devon and Somerset – environmental and stratigraphical implications (including discussion). Proceedings of the Ussher Society 4, 303–11.Google Scholar
Mayall, M. J. 1983. An earthquake origin for synsedimentary deformation in late Triassic (Rhaetian) lagoonal sequence, southwest Britain. Geological Magazine 120, 613–22.Google Scholar
Miliorizos, M. & Ruffell, A. 1998. Kinematics and geometry of the Watchet–Cothelstone–Hatch Fault System: implications for the structural history of the Wessex Basin and adjacent areas. In The Development, Evolution and Petroleum Geology of the Wessex Basin (ed. Underhill, J. R.), pp. 311–30. Geological Society of London, Special Publication no. 133.Google Scholar
Moore, C. 1881. On abnormal geological deposits in the Bristol District. Quarterly Journal of the Geological Society of London 37, 67–82.Google Scholar
Morawetz, S. 1965. Zur Frage der Dolinenverteilung und Dolinenbildung im Istrischen Karst. Petermann's Geographische Mitteilungen 109, 161–70.Google Scholar
Morbey, S. J. 1975. The palynostratigraphy of the Rhaetian Stage, Upper Triassic in the Kendalbachgraben, Austria. Palaeontographica Abteilung B 152, 1–75.Google Scholar
Morris, J. 1843. A catalogue of British fossils. London: British Museum (Natural History), 222 pp.Google Scholar
Mylroie, J. E. & Carew, J. I. 1995. Karst development on carbonate islands/Speleogenesis and Evolution of Karst Aquifers 1. In Unconformities in Carbonate Strata – Their Recognition and the Significance of Associated Porosity (eds Budd, D. A., Saller, A. H. & Harris, P. A.), pp. 55–76. American Association of Petroleum Geologists' Memoir no. 63.Google Scholar
Ohlson, M. & Tryterud, E. 2000. Interpretation of the charcoal record in forest soils: forest fires and their production and deposition of macroscopic charcoal. The Holocene 10, 519–25.Google Scholar
Olempska, E. 2004. Late Triassic spinocaudatan crustaceans from southwestern Poland. Acta Palaeontologia Polonica 49, 429–42.Google Scholar
Olsen, P. E., Kent, D. V., Sues, H.-D., Koeberl, C., Huber, H., Montanari, M., Rainforth, E. C., Powell, S. J., Szajna, M. J. & Hartline, B. W. 2002. Ascent of dinosaurs linked to an iridium anomaly at the Triassic–Jurassic boundary. Science 296, 1305–7.Google Scholar
Orbell, G. 1973. Palynology of the British Rhaeto-Liassic. Bulletin of the Geological Survey of Great Britain 44, 1–44.Google Scholar
Palmer, R. & Williams, D. 1984. Cave development under Andros Island, Bahamas. Transactions British Cave Research Association 11, 50–2.Google Scholar
Petersen, G. M. 1976. Pollen Analysis and the origin of Cave Sediments in the Central Kentucky Karst. Bulletin of the National Speleological Society 38, 53–8.Google Scholar
Rauhut, O. W. M. & Hungerbühler, A. 2000. A review of European Triassic theropods. GAIA 15, 75–88.Google Scholar
Rayfield, E. J., Barrett, P. M., McDonnell, R. A. & Willis, K. J. 2005. A geographical information system (GIS) study of Triassic vertebrate biochronology. Geological Magazine 142, 327–54.CrossRefGoogle Scholar
Renesto, S. 1995. A sphenodontid from the Norian (Late Triassic) of Lombardy (Northern Italy): a preliminary note. Modern Geology 20, 149–58.Google Scholar
Renesto, S. & Fraser, N. C. 2003. Drepanosaurid (Reptilia: Diapsida) remains from a Late Triassic fissure infilling at Cromhall Quarry (Avon, Great Britain). Journal of Vertebrate Paleontology 23, 703–5.CrossRefGoogle Scholar
Reynolds, S. H. 1938. A section of Rhaetic and associated strata at Chipping Sodbury, Glos. Geological Magazine 75, 97–103.CrossRefGoogle Scholar
Richardson, L. 1911. The Rhaetic and Contiguous Deposits of West, Mid and part of East Somerset. Quarterly Journal of the Geological Society of London 67, 1–74.CrossRefGoogle Scholar
Riley, H. & Stutchbury, S. 1836. A description of various fossil remains of three distinct saurian animals discovered in the autumn of 1834 in the Magnesian Conglomerate on Durdham Down near Bristol. Proceedings of the Geological Society 2, 397–9.Google Scholar
Riley, H. & Stutchbury, S. 1840. A description of various fossil remains of three distinct saurian animals recently discovered in the Magnesian Conglomerate near Bristol. Transactions of the Geological Society of London 5, 349–57.Google Scholar
Rixon, A. E. 1976. Fossil Animal Remains. Their preparation and conservation. London: Athlone Press, 304 pp.Google Scholar
Robinson, P. L. 1957 a. The Mesozoic fissures of the Bristol Channel area and their vertebrate faunas. Journal of the Linnean Society (Zoology) 43, 260–82.Google Scholar
Robinson, P. L. 1957 b. An unusual sauropsid dentition. Journal of the Linnean Society s(Zoology) 43, 283–93.Google Scholar
Robinson, P. L. 1962. Gliding lizards from the Upper Keuper of Great Britain. Proceedings of the Geological Society of London 1061, 137–46.Google Scholar
Ruffell, A. & Shelton, R. 1999. The control of sedimentary facies by climate during phases of crustal extension: examples from the Triassic of onshore and offshore England and Northern Ireland. Journal of the Geological Society, London 156, 779–89.Google Scholar
Säila, L. K. 2005. A new species of the sphenodontian reptile Clevosaurus from the Lower Jurassic of South Wales. Palaeontology 48, 817–31.Google Scholar
Savage, R. J. G. 1977. The Mesozoic strata of the Mendip Hills. In Geological excursions in the Bristol District (ed. Savage, R. J. G.), pp. 85–100. University of Bristol.Google Scholar
Savage, R. J. G. 1993. Vertebrate fissure faunas with special reference to Bristol Channel Mesozoic faunas. Journal of the Geological Society, London 150, 1025–34.CrossRefGoogle Scholar
Schoener, T. W. & Schoener, A. 1980. Densities, sex ratios and population structures in four species of Bahamian Anolis lizards. Journal of Animal Ecology 49, 19–53.CrossRefGoogle Scholar
Schuurman, W. M. L. 1977. Aspects of Late Triassic palysnology. 2. Palynology of the “Grès et Schiste à Avicula contorta” and “Argiles de Levallois” (Rhaetian) of north-eastern France and southern Luxembourg. Review of Palaeobotany & Palynology 23, 159–253.CrossRefGoogle Scholar
Schuurman, W. M. L. 1979. Aspects of Late Triassic palynology. 3. Palynology of latest Triassic and earliest Jurassic deposits of the northern limestone Alps in Austria and Southern Germany, with special reference to a palynological characterization of the Rhaetian Stage in Europe. Review of Palaeobotany and Palynology 27, 53–75.CrossRefGoogle Scholar
Seeley, H. G. 1891. On Agrosaurus mcgillivrayi, a saurischian reptile from the north east coast of Australia. Quarterly Journal of the Geological Society of London 47, 164–5.CrossRefGoogle Scholar
Seeley, H. G. 1895. On Thecodontosaurus and Palaeosaurus. Annals Magazine of Natural History 15, 144–63.Google Scholar
Shubin, N. H., Olsen, P. E. & Sues, H.-D. 1994. Early Jurassic small tetrapods from the McCoy Brook Formation of Nova Scotia, Canada. In In the shadow of the dinosaurs (eds Fraser, N. C. & Sues, H.-D.), pp. 242–50. New York: Cambridge University Press.Google Scholar
Shubin, N. H. & Sues, H.-D. 1991. Biogeography of early Mesozoic continental tetrapods: patterns and implications. Paleobiology 17, 214–30.CrossRefGoogle Scholar
Simms, M. J. 1990. Triassic Palaeokarst in Britain. Cave Science 17, 93–105.Google Scholar
Simms, M. J. 2006. Uniquely extensive soft-sediment deformation in the Rhaetian of the UK: Evidence for earthquake or impact? Palaeogeography, Palaeoclimatology, Palaeoecology 244, 407–23.CrossRefGoogle Scholar
Simms, M. J. & Ruffell, A. H. 1990. Climate and biotic change in the late Triassic. Journal of the Geological Society, London 147, 321–7.CrossRefGoogle Scholar
Simms, M. J., Ruffell, A. H. & Johnston, A. L. A. 1994. Biotic and climatic changes in the Carnian (Triassic) of Europe and adjacent areas. In In the shadow of the dinosaurs (eds Fraser, N. C. & Sues, H.-D.), pp. 352–65. New York: Cambridge University Press.Google Scholar
Skog, J. E. 2005. Report of the Committee for Fossil Plants 6. Taxon 54, 827.CrossRefGoogle Scholar
Smart, P. L., Dawans, J. M. & Whitaker, F. 1988. Carbonate dissolution in a modern mixing zone. Nature 335, 811–13.CrossRefGoogle Scholar
Strahan, A. & Cantrill, T. C. 1904. The geology of the South Wales Coalfield. Part VI. The country around Bridgend. Memoir, Geological Survey UK, 121 pp.Google Scholar
Sues, H.-D., Clark, J. C. & Jenkins, F. A. Jr. 1994. A review of the early Jurassic tetrapods from the Glen Canyon Group of the American SouthWest. In In the shadow of the dinosaurs (eds Fraser, N. C. & Sues, H.-D.), pp. 284–94. New York: Cambridge University Press.Google Scholar
Sues, H.-D. & Olsen, P. E. 1990. Triassic vertebrates of Gondwanan aspect from the Richmond Basin of Virginia. Science 249, 1020–3.CrossRefGoogle ScholarPubMed
Sues, H.-D., Olsen, P. E., Carter, J. G. & Scott, D. M. 2003. A new crocodylomorph archosaur from the Upper Triassic of North Carolina. Journal of Vertebrate Paleontology 23 (2), 329–43.Google Scholar
Sues, H. D. & Reisz, R. D. 1995. First record of the early Mesozoic sphenodontian Clevosaurus (Lepidosauria: Rhynchocephalia) from the Southern Hemisphere. Journal of Paleontology 69, 123–6.CrossRefGoogle Scholar
Sues, H.-D., Shubin, N. H. & Olsen, P. E. 1994. A new sphenodontian (Lepidosauria: Rhynchocephalia) from the McCoy Brook Formation (Lower Jurassic) of Nova Scotia, Canada. Journal of Vertebrate Paleontology 14, 327–40.Google Scholar
Sweeting, M. M. 1972. Karst Landforms. London: Macmillan, 362 pp.Google Scholar
Swift, A. & Martill, D. M. 1999. Fossils of the Rhaetian Penarth Group. London: The Palaeontological Association, 312 pp.Google Scholar
Swinton, W. E. 1939. A new Triassic Rhynocephalian from Gloucestershire. Annals Magazine Natural History Series II 4, 591–4.Google Scholar
Tarlo, L. B. H. 1962. Ancient animals of the uplands. New Scientist 15, 32–4.Google Scholar
Taylor, J. D., Braithwaite, C. J. R., Peake, J. F. & Arnold, E. N. 1979. Terrestrial faunas and habitats of Aldabra during the late Pleistocene. Philosophical Transactions of the Royal Society B 286, 47–66.Google Scholar
Thomas, J. B., Marshall, J. E. A., Mann, A. L., Summons, R. E. & Maxwell, J. R. 1993. Dinosteranes (4,23,24-trimethylsteranes) and other biological markers in dinoflagellate-rich, marine sediments of Rhaetian age. Organic Geochemistry 20, 91–104.Google Scholar
Torsvik, T. H., Van Der Voo, R., Meert, J. G., Mosar, J. & Walderhaug, H. J. 2001. Reconstructions of the continents around the North Atlantic at about the 60th parallel. Earth and Planetary Science Letters 187, 55–69.Google Scholar
Traverse, A. 2004. (1643) Proposal to conserve the fossil pollen morphogeneric name Classopollis against Corollina and Circulina. Taxon 53, 847–8.Google Scholar
Tucker, M. E. 1977. The marginal Triassic deposits of South Wales: continental facies and palaeogeography. Geological Journal 12, 169–88.Google Scholar
Turner, F. B. 1977. The dynamics of populations of squamates, crocodilians and rhynchocephalians. In Biology of the Reptilia 7 (eds Gans, C. & Tinkle, D. W.), pp. 157–264. London: Academic Press.Google Scholar
Tuweni, A. O. & Tyson, R. V. 1994. Organic facies variations in the Westbury Formation (Rhaetic, Bristol Channel, SW England). Organic Geochemistry 21, 1001–14.CrossRefGoogle Scholar
Walkden, G. M. & Fraser, N. C. 1993. Late Triassic fissure sediments and vertebrate faunas: Environmental change and faunal succession at Cromhall, South West Britain. Modern Geology 18, 511–35.Google Scholar
Walker, A. D. 1969. The reptile fauna of the lower Keuper Sandstone. Geological Magazine 106, 470–6.Google Scholar
Wall, G. R. T. & Jenkyns, H. C. 2004. The age, origin and tectonic significance of Mesozoic sediment-filled fissures in the Mendip Hills (SW England): implications for extension models and Jurassic sea-level curves. Geological Magazine 141, 471–504.CrossRefGoogle Scholar
Walsh, P. T., Boulter, M. C., Ijtaba, M. & Urbani, D. M. 1972. The preservation of the Neogene Brassington Formation of the Southern Pennines and its bearing on the evolution of Upland Britain. Journal of the Geological Society, London 128, 519–59.Google Scholar
Warrington, G. 1963. The occurrence of the branchiopod crustacean Euestheria in the Keuper Sandstone of Alderley Edge, Cheshire. Liverpool and Manchester Geological Journal 3, 315–19.CrossRefGoogle Scholar
Warrington, G. 1977. Palynological examination of Triassic (Keuper Marl and Rhaetic) deposits north-east and east of Bristol. Proceedings of the Ussher Society 4, 76–81.Google Scholar
Warrington, G. 1981. The indigenous micropalaeontology of British Triassic shelf sea deposits. In Microfossils from Recent and fossil shelf seas (eds Neale, J. W. & Brasier, M. D.), pp. 61–70. Chichester: Ellis Horwood.Google Scholar
Warrington, G. 1984. Late Triassic palynomorph records from Somerset. Proceedings of the Ussher Society 6, 29–34.Google Scholar
Warrington, G. 1996. Triassic spores and pollen. In Palynology: principles and applications (eds Jansonius, J. & McGregor, D. C.), pp. 755–66. American Association of Stratigraphic Palynologists Foundation 2.Google Scholar
Warrington, G. 2005. Late Triassic (Rhaetian) palynomorphs from near York, North Yorkshire, UK. Proceedings of the Yorkshire Geological Society 55, 305–6.CrossRefGoogle Scholar
Warrington, G. & Ivimey-Cook, H. C. 1992. Triassic. In Atlas of Palaeogeography and Lithofacies (eds Cope, J. C. W., Ingham, J. K. & Rawson, P. F.), pp. 97–106. Geological Society of London, Memoir no. 13.Google Scholar
Warrington, G. W. & Ivimey-Cook, H. C. 1995. The Late Triassic and Early Jurassic of coastal sections in west Somerset and South and Mid-Glamorgan. In Field Geology of the British Jurassic (ed. Taylor, P. D.), pp. 9–30. London: Geological Society.Google Scholar
Warrington, G., Ivimey-Cook, H. C., Edwards, R. A. & Whittaker, A. 1995. The Late Triassic–Early Jurassic succession at Selworthy, West Somerset, England. Proceedings of the Ussher Society 8, 426–32.Google Scholar
Warrington, G. & Whittaker, A. 1984. The Blue Anchor Formation (late Triassic) in Somerset. Proceedings of the Ussher Society 6, 100–7.Google Scholar
Warrington, G., Whittaker, A. & Scrivener, R. C. 1986. The late Triassic succession in central and eastern Somerset. Proceedings of the Ussher Society 6, 368–74.Google Scholar
Warrington, G. & Williams, B. J. 1984. The North Curry Sandstone Member (Late Triassic) near Taunton, Somerset. Proceedings of the Ussher Society 6, 82–7.Google Scholar
Wendt, J. 1971. Genese und fauna submariner sedimentorer spaltenfullungen im Mediterranen Jura. Palaeontographica Abteilung A 136, 122–92.Google Scholar
Whiteside, D. I. 1984. Fissure animals of Bristol and South Wales. Nature 309, 410.Google Scholar
Whiteside, D. I. 1986. The head skeleton of the Rhaetian sphenodontid Diphydontosaurus avonis gen. et. sp. nov. and the modernizing of a living fossil. Philosophical Transactions of the Royal Society, London B 312, 379–430.Google Scholar
Whiteside, D. I. & Marshall, J. E. A. 1985. Pre-Rhaetic therian mammals, comment to Fraser, Walkden & Stewart. Nature 318, 81–2.CrossRefGoogle Scholar
Whiteside, D. I. & Robinson, D. 1983. A glauconitic clay-mineral from a speleological deposit of Late Triassic age. Palaeogeography, Palaeoclimatology, Palaeoecology 4, 81–5.Google Scholar
Whittaker, R. J. 1998. Island Biogeography. Oxford University Press, 285 pp.Google Scholar
Wilford, G. E. 1964. The geology of Sarawak and Sabah Caves. Geological Survey Borneo region, Malaysia, Bulletin 6, 18.Google Scholar
Wings, O. 2004. Authigenic minerals in fossil bones from the Mesozoic of England: poor correlation with depositional environments. Palaeogeography, Palaeoclimatology, Palaeoecology 204, 15–32.Google Scholar
Wu, X.-C. 1994. Late Triassic-Early Jurassic sphenodontians from China and the phylogeny of the Sphenodontia. In In the shadow of the dinosaurs (eds Fraser, N. C. & Sues, H.-D.), pp. 38–69. New York: Cambridge University Press.Google Scholar
Yates, A. M. 2003. A new species of the primitive dinosaur Thecodontosaurus (Sarischia: Sauropodomorpha) and its implications for the systematics of early dinosaurs. Journal of Systematic Palaeontology 2, 1–42.CrossRefGoogle Scholar
Young, C. C. 1941 a. A complete osteology of Lufengosaurus huenei Young (gen. et sp. nov.). Palaeontologica Sinica (C) 7, 1–53.Google Scholar
Young, C. C. 1941 b. Gyposaurus sinensis (sp. nov.). a new Prosauropoda from the Upper Triassic Beds at Lufeng, Yunnan. Bulletin of the Geological Society China 21, 205–53.Google Scholar