An evolving pattern of fluvial dispersal responsible for the Lower Old Red Sandstone magnafacies has emerged from a stratigraphical and geographical consideration of palaeocurrent structures, clast lithology and fauna, sandstone composition, and sandstone detrital heavy-mineral assemblages. The evidence suggests that the Post-Ludlow (Downton) and lower Gedinnian rocks formed mainly in the joined Dingle–Shannon and Anglo–Welsh basins, fed from metamorphic complexes in NW Britain or beyond. Contemporaneous sedimentation in the Caledonian basin was restricted to its northeastern end. Later in the Gedinnian, the Dingle–Shannon and Angle–Welsh basins appear to have become separated by the uplift of a broad and extensive belt of mainly Lower Palaeozoic sedimentary and volcanic rocks; included in this belt were outcrops of Post-Ludlow strata representing the earlier stream of metamorphic detritus. Granite intrusion, which reduced the density of and consequently uplifted the crust, occurred widely in the regions dividing and bordering these basins, and may be causally connected with their separation in the Gedinnian. The Caledonian basin appears to have remained a separate feature.

Two new genera and species of haplolepids (Chondrostei), Protohaplolepis scotica and Blairolepis loanheadensis, are described from the ironstones of Loanhead, Midlothian (Namurian A, E1); this represents the earliest known record of the family. Protohaplolepis scotica (= Traquair's nomen nudum “Eurylepis scoticus”) shows many features in common with Westoll's hypothetical haplolepid “common ancestor X”, and is possibly ancestral to the Haplolepis–Microhaplolepis line within the family. Blairolepis loanheadensis, known only from a skull roof, shows greater similarities to Parahaplolepis and Pyritocephalus, but differs from these genera in having separate dermopterotics and parietals. Two subfamilies, the Haplolepinae and the Parahaplolepinae, are suggested to reflect the distinct evolutionary lines within the family.

A new species, Parahaplolepis westolli, is also described from Newsham, Northumberland (Westphalian B). This species shows a close resemblance to the American species Parahaplolepis tuberculata from Linton, Ohio.

A brief discussion of the evolutionary history of the family is presented, along with a summary of its stratigraphical and geographical distribution.

Recent collections of Sawdonia ornata have been made from the Strathmore Group which crops out in the Midland Valley of Scotland. The sediments are Lower Devonian, probably Emsian, in age. New morphological features discovered include planar branching of vegetative and fertile axes, abaxial emergences close to branch points and H branching. Well preserved cuticles have permitted reinterpretation of the structure of stornata, rosettes (hair bases), papillae and spines, and striated cuticle is identified as S. ornata. The morphology of sporangia and in situ spores is described. Pyritised axes have revealed details of xylem shape, maturation, changes in the vicinity of branching, tracheidal structure and the arrangement of xylem and cortex in deformed axes.

For much of its length in eastern Finland, the contact between the late Archaean granitoid terrane and early Proterozoic supracrustal rocks is that of a depositional unconformity tectonically modified during the Svecokarelian orogeny. In the Kaavi district, this modification has been extensive, particularly because of thrusting which resulted in tectonic slices of basement being present as inliers within the cover rocks and cover rocks as outliers within the main basement outcrop. Both basement and cover have undergone repeated deformation and metamorphic recrystallisation reflected by the presence of multiple tectonic fabrics and sequential mineral growths. However, it is only locally in major zones of movement that the basement rocks of the Presvecokarelides show major modification as the result of the deformation which had such a profound effect on the cover rocks during the development of the Svecokarelides. Over large areas, the products of four phases of deformation (DI–IV) remain in the basement gneisses together with the products of basic and acidic intrusion whose positions in the deformational sequence can be established‥

In the cover rocks, early nappe emplacement (Outokumpu nappe) was followed by the development of isoclinal folds associated with prograde metamorphism (D1). Regionally expressed asymmetrical folds, overturned to the E, with attenuation of overturned limbs and thrusting, involving basement, followed (D2). Subsequent deformation related to wrench faulting (D2c) was followed by the development of more open folds associated with declining geotherms (D3, D4); F3–F4 interference structures are expressed as domes, some of which were a locus of granitoid emplacement (Maarianvaara granitoid suite). D5 and D6 are expressed as cleavages and fractures.

Correlation of the structural features in the Kaavi district with those in the Outokumpu and adjacent districts forms the basis both for establishing the regional disposition of basement and cover units and for determining the structural evolution of this part of the Svecokarelides.