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Revision of Erpetosuchus (Archosauria: Pseudosuchia) and new erpetosuchid material from the Late Triassic ‘Elgin Reptile’ fauna based on μCT scanning techniques

Published online by Cambridge University Press:  06 November 2020

Davide FOFFA Open the ORCID record for Davide FOFFA [Opens in a new window] ,
Richard J. BUTLER ,
Sterling J. NESBITT ,
Stig WALSH ,
Paul M. BARRETT ,
Stephen L. BRUSATTE  and
Nicholas C. FRASER
Davide FOFFA*
Affiliation:
Department of Natural Sciences, National Museums Scotland, Chambers Street, EdinburghEH1 1JF, UK
Richard J. BUTLER
Affiliation:
School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, BirminghamB15 2TT, UK
Sterling J. NESBITT
Affiliation:
Department of Geosciences, Virginia Tech, 4044 Derring Hall (MC0420) 926 West Campus Drive, Blacksburg 24061, Virginia, USA
Stig WALSH
Affiliation:
Department of Natural Sciences, National Museums Scotland, Chambers Street, EdinburghEH1 1JF, UK School of GeoSciences, University of Edinburgh, Grant Institute, Hutton Road, EdinburghEH9 3FE, UK
Paul M. BARRETT
Affiliation:
Department of Earth Sciences, Natural History Museum, Cromwell Road, LondonSW7 5BD, UK
Stephen L. BRUSATTE
Affiliation:
Department of Natural Sciences, National Museums Scotland, Chambers Street, EdinburghEH1 1JF, UK School of GeoSciences, University of Edinburgh, Grant Institute, Hutton Road, EdinburghEH9 3FE, UK
Nicholas C. FRASER
Affiliation:
Department of Natural Sciences, National Museums Scotland, Chambers Street, EdinburghEH1 1JF, UK School of GeoSciences, University of Edinburgh, Grant Institute, Hutton Road, EdinburghEH9 3FE, UK
*
*Corresponding author. Email: [email protected]
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Abstract

The Late Triassic fauna of the Lossiemouth Sandstone Formation (LSF) from the Elgin area, Scotland, has been pivotal in expanding our understanding of Triassic terrestrial tetrapods. Frustratingly, due to their odd preservation, interpretations of the Elgin Triassic specimens have relied on destructive moulding techniques, which only provide incomplete, and potentially distorted, information. Here, we show that micro-computed tomography (μCT) could revitalise the study of this important assemblage. We describe a long-neglected specimen that was originally identified as a pseudosuchian archosaur, Ornithosuchus woodwardi. μCT scans revealed dozens of bones belonging to at least two taxa: a small-bodied pseudosuchian and a specimen of the procolophonid Leptopleuron lacertinum. The pseudosuchian skeleton possesses a combination of characters that are unique to the clade Erpetosuchidae. As a basis for investigating the phylogenetic relationships of this new specimen, we reviewed the anatomy, taxonomy and systematics of other erpetosuchid specimens from the LSF (all previously referred to Erpetosuchus). Unfortunately, due to the differing representation of the skeleton in the available Erpetosuchus specimens, we cannot determine whether the erpetosuchid specimen we describe here belongs to Erpetosuchus granti (to which we show it is closely related) or if it represents a distinct new taxon. Nevertheless, our results shed light on rarely preserved details of erpetosuchid anatomy. Finally, the unanticipated new information extracted from both previously studied and neglected specimens suggests that fossil remains may be much more widely distributed in the Elgin quarries than previously recognised, and that the richness of the LSF might have been underestimated.

Keywords

anatomyErpetosuchidaeErpetosuchus grantiLeptopleuronsystematics
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Earth and Environmental Science Transactions of The Royal Society of Edinburgh , Volume 111 , Issue 4 , December 2020 , pp. 209 - 233
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Copyright © The Author(s) 2020. Published by Cambridge University Press on behalf of The Royal Society of Edinburgh

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