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Further analysis of a Late Jurassic dinosaur bone-bed from the Morrison Formation of Montana, USA, with a computed three-dimensional reconstruction

Published online by Cambridge University Press:  23 September 2013

Glenn W. Storrs
Affiliation:
Cincinnati Museum Center, 1301 Western Avenue, Cincinnati, OH 45203, USA. Email:[email protected]
Sara E. Oser
Affiliation:
Montana State University, Department of Earth Sciences, P.O Box 173480, Bozeman, MT 59717-3480, USA. Email: [email protected]
Mark Aull
Affiliation:
University of Cincinnati, College of Engineering and Applied Science, School of Aerospace Systems, 2850 Campus Way, Cincinnati, OH 45221-0070, USA. Email: [email protected]

Abstract

A Late Jurassic sauropod bone-bed comprising thousands of individual skeletal elements in the Morrison Formation of Montana, USA, is re-analysed to further elucidate its taphonomic history. New data gained from detailed contextual sampling is assembled and presented to test the hypothesis of a drought-induced, mass-mortality assemblage, and its subsequent depositional emplacement by a debris flow. Large dinosaur bone-beds are well-known in the Morrison, yet the Mother's Day Quarry site is unique for the formation in depicting an apparent debris flow deposit. Use of a computer program tailored to the data allows three-dimensional reconstruction of the bone-bed and recreates the palaeoslope of the deposit, adding support to the debris flow hypothesis. Such digital manipulation of field data may allow a deeper understanding of other bone-bed origination events.

Type
Articles
Copyright
Copyright © The Royal Society of Edinburgh 2013 

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References

8. References

Abler, W. L. 1985. Skulls in fossil material: One mechanism contributing to their rarity. Journal of Paleontology 59, 249–50.Google Scholar
Allison, P. A. & Briggs, D. E. G. 1991. Taphonomy of nonmineralized tissues. In Allison, P. A. & Briggs, D. E. G. (eds) Taphonomy: Releasing the Data Locked in the Fossil Record, 2570. New York: Plenum Press.Google Scholar
Bakker, R. T. 1996. The real Jurassic Park: Dinosaurs and habitats at Como Bluff, Wyoming. In Morales, M. (ed.) The Continental Jurassic. Museum of Northern Arizona Bulletin 60, 3549.Google Scholar
Barnes, F. A. & Lockley, M. G. 1994. Trackway evidence for social sauropods from the Morrison Formation, eastern Utah (USA). Gaia 10, 3742.Google Scholar
Barnes, R. F. W. 1982. A note on elephant mortality in Ruaha National Park, Tanzania. African Journal of Ecology 20, 137–40.Google Scholar
Barrett, P. M. & Upchurch, P. 2005. Sauropod diversity through time: Macroevolutionary and paleoecological implications. In Curry Rogers, K. A. & Wilson, J. A.The Sauropods: Evolution and Paleobiology, 125–56. Berkeley: University of California Press.Google Scholar
Behrensmeyer, A. K. 1982. Time resolution in fluvial vertebrate asssemblages. Paleobiology 8, 211–27.Google Scholar
Behrensmeyer, A. K. 1987. Miocene fluvial facies and vertebrate taphonomy in northern Pakistan. In Ethridge, F. G., Flores, R. M. & Harvey, M. D. (eds) Recent Developments in Fluvial Sedimentology. SEPM Special Publication 39, 169–76. Tulsa, Oklahoma: Society for Sedimentary Geology.Google Scholar
Behrensmeyer, A. K. 1988. Vertebrate preservation in fluvial channels. In Behrensmeyer, A. K. & Kidwell, S. M. (eds) Ecological and Evolutionary Implications of Taphonomic Processes. Palaeogeography, Palaeoclimatology, Palaeoecology 63, 183–99.Google Scholar
Benvenuti, M. & Martini, I. P. 2002. Analysis of terrestrial hyperconcentrated flows and their deposits. Special Publication of the International Association of Sedimentologists 32, 167–93.Google Scholar
Berner, R. A. 1968. Calcium carbonate concretions formed by the decomposition of organic matter. Science 159, 195–97.Google Scholar
Britt, B. B., Eberth, D. A., Sheetz, R. D., Greenhalgh, B. W. & Stadtman, K. L. 2009. Taphonomy of debris-flow hosted dinosaur bone-beds at Dalton Wells, Utah (Lower Cretaceous, Cedar Mountain Formation, USA). Palaeogeography, Palaeoclimatology, Palaeoecology 280, 122.Google Scholar
Chure, D. J., Fiorillo, A. R. & Jacobsen, A. 1998a. Prey bone utilization by predatory dinosaurs in the Late Jurassic of North America, with comments on prey bone utilization throughout the Mesozoic. Gaia 15, 227–32.Google Scholar
Chure, D. J., Litwin, R., Hasiotis, S., Evanoff, E. & Carpenter, K. 1998b. The fauna and flora of the Morrison Formation. Modern Geology 23, 507–37.Google Scholar
Clauss, M. 2011. Sauropod biology and the evolution of gigantism. In Klein, N, Remes, K.Gee, C. T. & Sander, P. M. (eds) Biology of the Sauropod Dinosaurs: Understanding the Life of Giants, 37. Bloomington: Indiana University Press.Google Scholar
Conybeare, A. & Haynes, G. 1984. Observations on elephant mortality and bones in water holes. Quaternary Research 22, 189200.Google Scholar
Cope, E. D. 1877. On Amphicoelias, a genus of saurian from the Dakota epoch of Colorado. Proceedings of the American Philosophical Society 17, 242–46.Google Scholar
Corfield, T. F. 1973. Elephant mortality in Tsavo National Park, Kenya. East African Wildlife Journal 11, 339–68.Google Scholar
Craig, L. C., Holmes, C. N., Cadigan, R. A., Freeman, L., Mullers, T. E. & Weir, G. W. 1955. Stratigraphy of the Morrison and related formations, Colorado Plateau Region – A preliminary report. U.S. Geological Survey Bulletin 1009-E, 1168.Google Scholar
Dasgupta, P. 2003. Sediment gravity flow – the conceptual problems. Earth-Science Reviews 62, 265–81.Google Scholar
Day, J. T., Upchurch, P., Norman, D. B., Gale, A. S. & Powell, H. P. 2002. Sauropod trackways, evolution, and behavior. Science 296, 1659.Google Scholar
Demko, T. M., Currie, B. S. & Nicoll, K. A. 2004. Regional paleoclimatic and stratigraphic implications of paleosols and fluvial/overbank architecture in the Morrison Formation (Upper Jurassic), Western Interior, USA. Sedimentary Geology 167, 115–35.Google Scholar
Demko, T. M. & Parrish, J. T. 1998. Paleoclimatic setting of the Upper Jurassic Morrison Formation. Modern Geology 22, 283–96.Google Scholar
Dodson, P., Behrensmeyer, A. K., Bakker, R. T. & McIntosh, J. S. 1980. Taphonomy and paleoecology of the dinosaur beds of the Jurassic Morrison Formation. Paleobiology 6, 208–32.Google Scholar
Dudley, J. P., Craig, G. C., Gibson, D. St. C., Haynes, G. & Klimowicz, J. 2001. Drought mortality of bush elephants in Hwange National Park, Zimbabwe. African Journal of Ecology 39, 187–94.Google Scholar
Dunagan, S. P. & Turner, C. E. 2004. Regional paleohydrologic and paleoclimatic settings of wetland/lacustrine depositional systems in the Morrison Formation (Upper Jurassic), Western Interior, USA. Sedimentary Geology 167, 269–96.Google Scholar
Eberth, D. A., Britt, B. B., Sheetz, R., Stadtman, K. L. & Brinkman, D. B. 2006. Dalton Wells: Geology and significance of debris-flow-hosted dinosaur bonebeds in the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah, USA. Palaeogeography, Palaeoclimatology, Palaeoecology 236, 217–45.Google Scholar
Eberth, D. A. & Getty, M. A. 2005. Ceratopsian bonebeds: Occurrence, origins, and significance. In Currie, P. J. & Koppelhus, E. B. (eds) Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed, 501–36. Bloomington: Indiana University Press.Google Scholar
Engelmann, G. F., Chure, D. J. & Fiorillo, A. R. 2004. The implications of a dry climate for the paleoecology of the fauna of the Upper Jurassic Morrison Formation. Sedimentary Geology 167, 297308.Google Scholar
Farlow, J. O., Pittman, J. G. & Hawthorne, J. M. 1989. Brontopodus birdi, Lower Cretaceous sauropod footprints from the US Gulf coastal plain. In Gillette, D. D. & Lockley, M. G. (eds) Dinosaur Tracks and Traces, 371–94. Cambridge, UK: Cambridge University Press.Google Scholar
Fastovsky, D. E., Clark, J. M., Strater, N. H., Montellano, M., Hernandez, R. & Hopson, J. A. 1995. Depositional environments of a Middle Jurassic terrestrial vertebrate assemblage, Huizachal Canyon, Mexico. Journal of Vertebrate Paleontology 15, 561–75.Google Scholar
Foster, J. R. 2003. Paleoecological analysis of the vertebrate fauna of the Morrison Formation (Upper Jurassic), Rocky Mountain Region, USA. Bulletin of the New Mexico Museum of Natural History and Science 23. Albuquerque: New Mexico Museum of Natural History and Science. 95 pp.Google Scholar
Foster, J. R. 2007. Jurassic West: The Dinosaurs of the Morrison Formation and Their World. Bloomington: Indiana University Press. 389 pp.Google Scholar
Gilmore, C. W. 1925. A nearly complete articulated skeleton of Camarasaurus, a saurischian dinosaur from the Dinosaur National Monument. Memoirs of the Carnegie Museum 10, 347–84.Google Scholar
Gilmore, C. W. 1936. Osteology of Apatosaurus with special reference to specimens in the Carnegie Museum. Memoirs of the Carnegie Museum 11, 175300.Google Scholar
Harris, J. D. 2006. The axial skeleton of the dinosaur Suuwassea emilieae (Sauropoda: Flagellicaudata) from the Upper Jurassic Morrison Formation of Montana, USA. Palaeontology 49, 1091–121.Google Scholar
Harris, J. D. 2007. The appendicular skeleton of Suuwassea emilieae (Sauropoda: Flagellicaudata) from the Upper Jurassic Morrison Formation of Montana (USA). Geobios 40, 501–22.Google Scholar
Harris, J. D. & Dodson, P. 2004. A new diplodocoid sauropod dinosaur from the Upper Jurassic Morrison Formation of Montana, USA. Paleontologica Polonica 49, 197210.Google Scholar
Hatcher, J. B. 1901. Diplodocus (Marsh): Its osteology, taxonomy, and probable habits, with a restoration of the skeleton. Memoirs of the Carnegie Museum 1, 163.Google Scholar
Haynes, G. 1985. Age profiles in elephant and mammoth bone assemblages. Quaternary Research 24, 333–45.Google Scholar
Haynes, G. 1987. Proboscidean die-offs and die-outs: Age profiles in fossil collections. Journal of Archaeological Science 14, 659–68.Google Scholar
Heinrich, W.-D. 1999. The taphonomy of dinosaurs from the Upper Jurassic of Tendaguru (Tanzania) based on field sketches of the German Tendaguru Expedition (1909–1913). Mitteilungen aus dem Museum für Naturkunde Berlin. Geowissenschaftliche Reihe 2, 2561.Google Scholar
Hill, A. & Behrensmeyer, A. K. 1984. Disarticulation patterns of some modern East African mammals. Paleobiology 10, 366–67.Google Scholar
Holland, W. 1906. The osteology of Diplodocus Marsh. Memoirs of the Carnegie Museum 2, 225–78.Google Scholar
Horner, J. R. & Dobb, E. 1997. Dinosaur Lives: Unearthing an Evolutionary Saga. New York: HarperCollins. 244 pp.Google Scholar
Hungerbühler, A. 1998. Taphonomy of the prosauropod dinosaur Sellosaurus, and its implications for carnivore faunas and feeding habits in the Late Triassic. Palaeogeography, Palaeoclimatology, Palaeoecology 143, 129.Google Scholar
Iverson, R. M., Reid, M. E. & LaHusen, R. G. 1997. Debris-flow mobilization from landslides. Annual Review of Earth and Planetary Sciences 25, 85138.Google Scholar
Jennings, D. S. & Hasiotis, S. T. 2006. Taphonomic analysis of a dinosaur feeding site using geographic information systems (GIS), Morrison Formation, southern Bighorn Basin, Wyoming, USA. Palaios 21, 480–92.Google Scholar
Jensen, J. A. 1985. Three new sauropod dinosaurs from the Upper Jurassic of Colorado. Great Basin Naturalist 45, 697709.Google Scholar
Kirkland, J. I., Carpenter, K., Hunt, A. P. & Scheetz, R. D. 1998. Ankylosaur (Dinosauria) specimens from the Upper Jurassic Morrison Formation. Modern Geology 22, 145–77.Google Scholar
Kohl, M. F. & McIntosh, J. S. 1997. Discovering Dinosaurs in the Old West – The Field Journals of Arthur Lakes. Washington, DC: Smithsonian Institution Press. 198 pp.Google Scholar
Kowallis, B. J., Christiansen, E. H. & Deino, A. L. 1991. Age of the Brushy Basin Member of the Morrison Formation, Colorado Plateau, western USA. Cretaceous Research 12, 483–93.Google Scholar
Kowallis, B. J., Christiansen, E. H., Deino, A. L., Peterson, F., Turner, C. E., Kunk, M. J. & Obradovich, J. D. 1998. The age of the Morrison Formation. Modern Geology 22, 235–60.Google Scholar
Lauters, P., Bolotsky, Y. L., Van Itterbeeck, J. & Godefroit, P. 2008. Taphonomy and age profile of a latest Cretaceous dinosaur bone bed in far eastern Russia. Palaios 23, 153–62.Google Scholar
Lockley, M. G. 1991. Tracking Dinosaurs. Cambridge, UK: Cambridge University Press. 238 pp.Google Scholar
Lockley, M. G., Houck, K. J. & Prince, N. K. 1986. North America's largest dinosaur trackway site: implications for Morrison paleoecology. Geological Society of America Bulletin 97, 1163–76.Google Scholar
Lockley, M. G., Foster, J. R. & Hunt, A. P. 1998. A short summary of dinosaur tracks and other fossil footprints from the Morrison Formation. Modern Geology 22, 277–90Google Scholar
Lockley, M. G., Wright, J., White, D., Matsukawa, M., Jianjun, L., Lu, F. & Hong, L. 2002. The first sauropod trackways from China. Cretaceous Research 23, 363–81.Google Scholar
Lovelace, D. M., Hartman, S. A. & Wahl, W. R. 2007. Morphology of a specimen of Supersaurus (Dinosauria, Sauropoda) from the Morrison Formation of Wyoming, and a re-evaluation of diplodocid phylogeny. Arquivos do Museu Nacional, Rio de Janeiro 65, 527–44.Google Scholar
Lucas, S. G. & Kirkland, J. I. 1998. Preliminary report on Conchostraca from the Upper Jurassic Morrison Formation, western United States. Modern Geology 22, 415–22.Google Scholar
Lull, R. S. 1919. The sauropod dinosaur Barosaurus Marsh. Memoirs of the Connecticut Academy of Arts and Science 6, 142.Google Scholar
Madsen, J. H. Jr. 1976. Allosaurus fragilis – a revised osteology. Utah Geological and Mineral Survey Bulletin 109, 1163.Google Scholar
Madsen, J. H. Jr., McIntosh, J. S. & Berman, D. S. 1995. Skull and atlas-axis complex of the Upper Jurassic sauropod Camarasaurus Cope (Reptilia: Saurischia). Bulletin of the Carnegie Museum of Natural History 31, 1115.Google Scholar
Madsen, J. H. Jr. & Welles, S. P. 2000. Ceratosaurus (Dinosauria, Theropoda), a revised osteology. Utah Geological Survey Miscellaneous Publication 00-2, 180.Google Scholar
Manley, K. 1993. Surface polish measurements from bona fide and suspected sauropod dinosaur gastroliths, wave and stream transported clasts. Ichnos 2, 167–69.Google Scholar
Marsh, O. C. 1877. Notice of new dinosaurian reptiles from the Jurassic formation. American Journal of Science and Arts 14, 514–16.Google Scholar
Marsh, O. C. 1878. Principal characters of American Jurassic dinosaurs. Part 1. American Journal of Science 3, 411–16.Google Scholar
Marsh, O. C. 1890. Description of new dinosaurian reptiles. American Journal of Science 39, 8186.Google Scholar
Marsh, O. C. 1896. The Dinosaurs of North America. Annual Report of the United States Geological Survey 16. 282 pp.Google Scholar
Mateus, O. 2006. Late Jurassic dinosaurs from the Morrison Formation, the Lourinhã and Alcobaça formations (Portugal), and the Tendaguru Beds (Tanzania): a comparison. In Foster, J. R. & Lucas, S. G. (eds.) Paleontology and Geology of the Upper Jurassic Morrison Formation. New Mexico Museum of Natural History and Science Bulletin 36, 223–31. Albuquerque: New Mexico Museum of Natural History and Science. 249 pp.Google Scholar
McIntosh, J. S. 1981. Annotated catalogue of the dinosaurs (Reptilia, Archosauria) in the collections of the Carnegie Museum of Natural History. Bulletin of the Carnegie Museum of Natural History 18, 167.Google Scholar
McIntosh, J. S. 1990. Sauropoda. In Weishampel, D. B., Dodson, P. & Osmólska, H. (eds) The Dinosauria, 345401. Berkeley: University of California Press.Google Scholar
McIntosh, J. S. 2005. The genus Barosaurus Marsh (Sauropoda, Diplodocidae). In Tidwell, V. & Carpenter, K. (eds) Thunder Lizards: The Sauropodomorph Dinosaurs, 3877. Bloomington: Indiana University Press.Google Scholar
McIntosh, J. S. & Carpenter, K. 1998. The holotype of Diplodocus longus, with comments on other specimens of the genus. Modern Geology 23, 85110.Google Scholar
McNab, B. K. 2009. Resources and energetics determined dinosaur maximal size. Proceedings of the National Academy of Science 106, 12184–88.Google Scholar
Monaco, P. E. 1998. A short history of dinosaur collecting in the Garden Park Fossil Area, Cañon City, Colorado. Modern Geology 23, 465–80.Google Scholar
Munsell Color. 2009. Geological Rock-Color Chart. 4300 44th Street, Grand Rapids, Michigan 49512.Google Scholar
Myers, T. S. & Fiorillo, A. 2009. Evidence for gregarious behavior and age segregation in sauropod dinosaurs. Palaeogeography, Palaeoclimatology, Palaeoecology 274, 96104.Google Scholar
Myers, T. S. & Storrs, G. W. 2007. Taphonomy of the Mother's Day Quarry, Upper Jurassic Morrison Formation, south-central Montana, USA. Palaios 22, 652–66.Google Scholar
Oba, G. 2001. The effect of multiple droughts on cattle in Obbu, northern Kenya. Journal of Arid Environments 49, 375–86.Google Scholar
Osborn, H. F. & Mook, C. C. 1921. Camarasaurus, Amphicoelias, and other sauropods of Cope. Memoirs of the American Museum of Natural History 3, 249386.Google Scholar
Parrish, J. T., Peterson, F. & Turner, C. E. 2004. Jurassic “savannah” – plant taphonomy and climate of the Morrison Formation (Upper Jurassic, Western USA). Sedimentary Geology 167, 137–62.Google Scholar
Peterson, F. 1988. Stratigraphy and nomenclature of Middle and Upper Jurassic rocks, Western Colorado Plateau, Utah and Arizona. United States Geological Survey Bulletin 1633-B, 1756.Google Scholar
Peterson, F. & Turner, C. E. 1998. Stratigraphy of the Ralston Creek and Morrison formations (Upper Jurassic) near Denver, Colorado. Modern Geology 22, 338.Google Scholar
Rao, J. S. 1972. Some variants of chi-square for test uniformity on the circle. Zeitschrift für Wahrscheinlichkeitstheorie und verwandte gebiete 22, 3344.Google Scholar
Rao, J. S. 1976. Some tests based on the arc-lengths for the circle. Sankhya: The Indian Journal of Statistics 38, 329–38.Google Scholar
Richmond, D. R. & Morris, T. H. 1998. Stratigraphy & cataclysmic deposition of the Dry Mesa Dinosaur Quarry, Mesa County, Colorado. Modern Geology 22, 121–43.Google Scholar
Riggs, E. S. 1903. Structure and relationships of opisthocoelian dinosaurs. Part I. Apatosaurus Marsh. Publications of the Field Columbian Museum, Geological Series 2, 165–96.Google Scholar
Rogers, R. R. 1990. Taphonomy of three dinosaur bone beds in the Upper Cretaceous Two Medicine Formation of northwestern Montana: evidence for drought-related mortality. Palaios 5, 394413.Google Scholar
Rogers, R. R. 2005. Fine-grained debris flows and extraordinary vertebrate burials in the Late Cretaceous of Madagascar. Geology 33, 297300.Google Scholar
Russell, G. S. & Levitin, D. J. 1995. An expanded table of probability values for Rao's Spacing Test. Communications in Statistics: Simulation and Computation 24, 879–88.Google Scholar
Sander, P. M. & Clauss, M. 2008. Sauropod gigantism. Science 322, 200–01.Google Scholar
Sanders, F., Manley, K. & Carpenter, K. 2001. Gastroliths from the Lower Cretaceous sauropod Cedarosaurus weiskopfae. In Tanke, D. & Carpenter, K. (eds) Mesozoic Vertebrate Life, 166–80. Bloomington: Indiana University Press.Google Scholar
Steiner, M. B. 1998. Age, correlation and tectonic implications of Morrison Formation paleomagnetic data, including rotation of the Colorado Plateau. Modern Geology 22, 261–81.Google Scholar
Stokes, W. L. 1944. Morrison Formation and related deposits in and adjacent to the Colorado Plateau. Geological Society of America Bulletin 55, 951–92.Google Scholar
Stovall, J. W. 1938. The Morrison of Oklahoma and its dinosaurs. Journal of Geology 46, 583600.Google Scholar
Tasch, P. 1987. Fossil Conchostraca of the southern hemisphere and continental drift. Geological Society of America Memoir 165, 1290.Google Scholar
Turner, C. E. & Peterson, F. 1999. Biostratigraphy of dinosaurs in the Upper Jurassic Morrison Formation of the Western Interior, U.S.A. Utah Geological Survey Miscellaneous Publication 99-1, 77114.Google Scholar
Turner, C. E. & Peterson, F. 2004. Reconstruction of the Upper Jurassic Morrison Formation extinct ecosystem – A synthesis. Sedimentary Geology 167, 309–55.Google Scholar
Upchurch, P., Barrett, P. M. & Dodson, P. 2004a. Sauropoda. In Weishampel, D. B., Dodson, P. & Osmólska, H. (eds) The Dinosauria (2nd edition), 259322. Berkeley: University of California Press.Google Scholar
Upchurch, P., Tomida, Y. & Barrett, P. M. 2004b. A new specimen of Apatosaurus ajax (Sauropoda: Diplodocidae) from the Morrison Formation (Upper Jurassic) of Wyoming, USA. National Science Museum Monographs 26, 1108.Google Scholar
Vallance, J. W. & Scott, K. M. 1997. The Osceola Mudflow from Mount Rainier: Sedimentology and hazard implications of a huge clay-rich debris flow. Geological Society of America Bulletin 109, 143–63.Google Scholar
Van Itterbeeck, J., Bolotsky, Y. L., Bultynck, P. & Godefroit, P. 2005. Stratigraphy, sedimentology and palaeoecology of the dinosaur-bearing Kundur section (Zeya-Bureya Basin, Amur Region, far eastern Russia). Geological Magazine 142, 735–50.Google Scholar
Voorhies, M. R. 1969. Taphonomy and population dynamics of an early Pliocene vertebrate fauna, Knox County, Nebraska. University of Wyoming Contributions to Geology Special Paper 1, 169.Google Scholar
Webb, J. A. 1979. A reappraisal of the paleoecology of conchostracans (Crustacea: Branchiopoda). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 158, 259–79.Google Scholar
Weigelt, J. 1989. Recent Vertebrate Carcasses and Their Paleobiological Implications. Chicago: University of Chicago Press. 188 pp.Google Scholar
Wilhite, D. R. 2005. Variation in the appendicular skeleton of North American sauropod dinosaurs: taxonomic implications. In Tidwell, V. & Carpenter, K. (eds) Thunder Lizards: The Sauropodomorph Dinosaurs, 268301. Bloomington: Indiana University Press.Google Scholar
Wilson, J. A. 2002. Sauropod dinosaur phylogeny: Critique and cladistic analysis. Zoological Journal of the Linnean Society 136, 217–76.Google Scholar
Wright, J. L. 2005. Steps in understanding sauropod biology: The importance of sauropod tracks. In Curry Rogers, K. A. & Wilson, J. A. (eds) The Sauropods – Evolution and Paleobiology, 252–84. Berkeley: University of California Press.Google Scholar
Zaleha, M. J. & Wiesemann, S. A. 2005. Hyperconcentrated flows and gastroliths: sedimentology of diamictites and wackes of the upper Cloverly Formation, Lower Cretaceous, Wyoming, U.S.A. Journal of Sedimentary Research 75, 4354.Google Scholar
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