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A REVISION OF THE NEARCTIC SPECIES OF THE TACHYDROMIINE FLY GENUS STILPON LOEW (DIPTERA: EMPIDOIDEA)

Published online by Cambridge University Press:  31 May 2012

J.M. Cumming  and
B.E. Cooper
J.M. Cumming
Affiliation:
Biological Resources Division, Centre for Land and Biological Resources Research, Agriculture Canada, Research Branch, Central Experimental Farm, Ottawa, Ontario, Canada K1A 0C6
B.E. Cooper
Affiliation:
Biological Resources Division, Centre for Land and Biological Resources Research, Agriculture Canada, Research Branch, Central Experimental Farm, Ottawa, Ontario, Canada K1A 0C6
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Abstract

This revision recognizes 13 species of Stilpon Loew in the Nearctic Region, including nine new species. A key is provided for the identification of adult specimens, all species are described, and their geographical distributions are mapped. Two informal species groups are recognized in the Nearctic Region, with the following included species: (1) S. graminum group — S. campestris Cumming sp.nov. (type-locality Assiniboia, Saskatchewan, Canada); (2) S. varipes group — S. chillcotti Cumming sp.nov. (type-locality 10 km E Evergreen, Alabama, USA), S. ctenistes Cumming sp.nov. (type-locality Salmon Creek, New York, USA), S. curvipes Melander, S. limitaris Cumming sp.nov. (type-locality Sapelo Island,Georgia,USA), S. pauciseta Melander, S. pilomus Cumming sp.nov. (type-locality Franconia, New Hampshire, USA), S. spinipes Melander, S. tribulosus Cumming sp.nov. (type-locality Gatlinburg, Tennessee, USA), S. tyconyx Cumming sp.nov. (type-locality Bedford, Massachusetts, USA), S. varipes Loew, S. vockerothi Cumming sp.nov. (type-locality Ottawa, Ontario, Canada), and S. wirthi Cumming sp.nov. (type-locality Dennisport, Massachusetts, USA). A world list of described species of Stilpon, including synonyms, is provided. Stilpon pectiniger Melander is a new junior synonym of S. varipes Loew and S. demnatensis Vaillant is indicated as a nomen nudum. Stilpon pleuriticus Melander, previously considered to belong within Stilpon, is excluded from the genus, and S. obscuripes Adams is transferred as Crossopalpus obscuripes (Adams) comb.nov. Lectotypes are designated for S. pauciseta Melander and S. pectiniger Melander. Homologies of previously confused structures of the male and female terminalia are summarized with reference to other Empidoidea. The monophyly of Stilpon is justified, including discussion of the phylogenetic relationships of the genus to the remainder of the Tachydromiinae. All described species of Stilpon, including extralimital species, are assigned to one of three proposed informal species groups (S. divergens, S. graminum, or S. varipes group), and zoogeographic patterns are discussed.

Résumé

Cette révision reconnaît 13 espèces de Stilpon Loew dans la région néarctique, dont neuf espèces inédites. On trouvera ici une clé d’identification des adultes, une description de toutes les espèces ainsi que des cartes de leurs répartitions géographiques. Deux groupes informels d’espèces sont reconnus dans la région néarctique, parmi lesquels on retrouve les espèces suivantes : (1) dans le groupe S. graminum — S. campestris Cumming sp.nov. (localité type, Assiniboia, Saskatchewan, Canada); (2) dans le groupe S. varipes — S. chillcotti Cumming sp.nov. (localité type, 10 km à l’est d’Evergreen, Alabama, É.-U.), S. ctenistes Cumming sp.nov. (localité type, Salmon Creek, New York, É.-U.), S. curvipes Melander, S. limitaris Cumming sp.nov. (localité type, Sapelo Island, Géorgie, É.-U.), S. pauciseta Melander, S. pilomus Cumming sp.nov. (localité type, Franconia, New Hampshire, É.-U.), S. spinipes Melander, S. tribulosus Cumming sp.nov. (localité type, Gatlinburg, Tennessee, É.-U.), S. tyconyx Cumming sp.nov. (localité type, Bedford, Massachusetts, É.-U.), S. varipes Loew, S. vockerothi Cumming sp.nov. (localité type, Ottawa, Ontario, Canada) et S. wirthi Cumming sp.nov. (localité type, Dennisport, Massachusetts, É.-U.). Une liste à l’échelle mondiale des espèces décrites de Stilpon, synonymes inclus, est proposée. Stilpon pectiniger Melander est un nouveau synonyme récent de S. varipes Loew et S. demnatensis Vaillant devient nomen nudum. Stilpon pleuriticus Melander, considéré précédemment comme un Stilpon, est exclus du genre et S. obscuripes Adams est transféré au genre Crossopalpus, devenant Crossopalpus obs-curipes (Adams) comb.nov. DeslectotypesdeS. pauciseta Melander et de S. pectiniger Melander ont été choisis. L’homologie de certaines structures des genitalia mâle et femelle, au sujet desquelles il y avait de la confusion, a été redéfinie par comparaison à d’autres Empidoidea. La monophylie de Stilpon est justifié et les relations phylogénétiques entre ce genre et les autres Tachydromiinae sont examinées. Toutes les espèces décrites de Stilpon, y compris les espèces existant hors des limites de l’étude, ont été classifiées dans l’un des trois groupes informels proposés (groupes S. divergens, S. graminum ou S. varipes) et leurs répartitions zoogéographiques font l’objet d’une discussion.

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The Canadian Entomologist , Volume 124 , Issue 6 , December 1992 , pp. 951 - 998
Copyright
Copyright © Entomological Society of Canada 1992

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