Quackgrass is a problematic agricultural weed in the temperate zones of the world and is difficult to control without herbicides or intensive tillage. However, it may be possible to control quackgrass with less environmental impact by combining multiple low-intensity control methods. A pot experiment was conducted in July to October 2012 and repeated in June to September 2013 to investigate the effect of rhizome fragmentation, competition from white clover, shoot-cutting frequency, and cutting height on quackgrass. Rhizome fragmentation was expected to result in more, but weaker, quackgrass shoots that would be more vulnerable to shoot cutting and competition. However, by 20 d past planting, rhizome fragmentation did not change the total number of quackgrass shoots per pot, because an increase in main shoots was offset by a decrease in tiller numbers. Rhizome fragmentation did not reduce quackgrass biomass acquisition during the experimental period. Although rhizome fragmentation did reduce total fructan content, it did not enhance the effect of clover competition, shoot-cutting frequency, or shoot-cutting height. Clover competition by itself reduced quackgrass shoot numbers by 72%, rhizome biomass by 81%, and belowground fructan concentration by 10 percentage points, compared with no competition. The more frequently quackgrass shoots were cut, the less biomass quackgrass acquired, and a high shoot-cutting frequency (each time quackgrass reached 2 leaves) resulted in a lower belowground fructan concentration than a low shoot-cutting frequency (at 8 leaves). However, in pots without competition, a higher shoot-cutting frequency resulted in more quackgrass shoots. A lower shoot-cutting height (25 mm) had more impact when shoot cutting was more frequent. In conclusion, rhizome fragmentation did not reduce the number of quackgrass shoots or rhizome biomass, but competition from white clover, a high shoot-cutting frequency, and a low shoot-cutting height strongly suppressed quackgrass biomass and fructan acquisition.

When grown without soybean [Glycine max (L.) Merr.] interference, eastern black nightshade (Solanum ptycanthum Dun.) planted in May produced as many as 7000 berries and 800000 seeds per plant. Eastern black nightshade planted in mid-July produced as many as 100 berries, but those planted in mid-August failed to produce berries. Eastern black nightshade produced less than 85 berries per plant when planted with soybeans in May, 0 to 3 berries per plant when planted into soybeans in June, and no berries when planted into soybeans in July. However, when soybeans were defoliated in July to simulate hail injury, eastern black nightshade that had been planted with soybeans in May produced up to 1600 berries per plant, and those planted into the defoliated soybeans in July produced up to 58 berries per plant. We suggest, therefore, that in vigorously growing soybeans, control of eastern black nightshade emerging in the row would only be required through June to prevent berry production. If, however, the soybean canopy were disturbed by hail or other factors, eastern black nightshade seedlings emerging in late July or early August might produce berries by soybean harvest.

The triethylamine salt of triclopyr [(3,5,6-trichloro-2-pyridinyl)oxy] acetic acid and the potassium salt of picloram (4-amino-3,5,6-trichloropicolinic acid) were applied as soil, foliar, and soil-plus-foliar treatments to greenhouse-grown huisache [Acacia farnesiana (L.) Willd.]. At 1.12 kg/ha, picloram was more effective (97 to 100% defoliation) than triclopyr (51 to 83%) in defoliating huisache. Triclopyr was more effective at 2.24 kg/ha than at 1.12 kg/ha in defoliating huisache, except when applied to the soil. Soil or soil-plus-foliar treatments of triclopyr at 2.24 kg/ha were not significantly different from those of picloram at 1.12 kg/ha 15 weeks after treatment. Triclopyr and picloram content was determined in and on leaves, stems, and roots at 0, 3, 10, and 30 days after treatment. More picloram than triclopyr was found in the leaves 30 days after treatment with foliar sprays at equal rates. Soil, foliar, and soil-plus-foliar treatments resulted in greater accumulation of picloram than triclopyr in stem tissue. Herbicide concentrations in roots were usually low. Reasons for the greater herbicidal activity of picloram than of triclopyr in huisache are not clear; however, the greater accumulation of picloram than of triclopyr in leaf and stem tissue may partially explain the difference in herbicide activity.

Enhancement of the canker causing ability of Botryosphaeria ribis on melaleuca was studied with respect to stress from simulated drought, low temperature, and defoliation treatments. Low xylem water potential was related to increased level of canker development and subsequent tree mortality. Canker development was enhanced by low temperature treatments with alternating exposure to 6 C for 3 d followed by 4 d at 30 (±5) C for 8 wk. Partial defoliation did not affect canker development but complete defoliation of B. ribis-inoculated ramets resulted in tree mortality within 4 wk. Callusing of melaleuca wounds was either reduced or prevented in stressed trees. These observations suggest that stress induced on the tree enhances the tree-killing efficacy of this fungus.

A laboratory study was conducted to determine the effects of two adjuvants and temperatures at time of treatment on efficacy, absorption, and translocation of thidiazuron defoliant on cotton. Five days after treatment at 30/21 C day/night temperatures, leaf drop was 17% with no adjuvant, 37% with addition of crop oil concentrate, 40% with ammonium sulfate, and 75% with two adjuvants combined. At 21/13 C day/night temperatures, there was less than 10% leaf drop with all treatments. At 10 d after treatment, leaf drop was not different among treatments at the high or low temperatures. Shoot regrowth at high and low temperature was reduced 55 to 60% with addition of both adjuvants and 44 to 50% with each adjuvant or with no adjuvant when compared to plants defoliated by hand. Absorption of 14C-thidiazuron was not affected by variations in temperature during the time of treatment but was affected by adjuvants. With no adjuvants, absorption was 7 to 10%. With 1.25% by vol crop oil concentrate, absorption was 33 to 46%. Addition of ammonium sulfate resulted in 18 to 19% absorption, and the combination of ammonium sulfate and crop oil concentrate increased absorption to 65 to 68%. There was no movement of radiolabel away from treated leaves as determined by autoradiographs of treated plants.

The combined influence of plant competition and defoliation on development of Canada thistle was investigated in a 2-yr field study. Plant competition was induced by seeding tall fescue and crown vetch. Artificial defoliation was used to simulate various levels of leaf removal by insects. Both defoliation and induced competition reduced biomass of Canada thistle but their impact varied with environmental conditions. Defoliation had a greater detrimental influence than induced competition on thistle biomass in a dry year when growth of the plant competitors was suppressed. In a wet year, induced competition was more important in suppressing Canada thistle than defoliation, and moderate levels of defoliation (25%), applied once when the thistles were 12 to 15 cm in diam, stimulated root weight within the top 20 cm of soil. Reduction in thistle biomass increased with level of defoliation and was greatest when defoliation was applied repeatedly at 14-d intervals in the presence of induced competition. Crown vetch showed very little growth in one season and tall fescue was the primary source of competition for the thistles. The results confirm the hypothesis that combined stresses can substantially reduce biomass development of Canada thistle plants.

Sulfur cinquefoil is an invasive, perennial forb on rangelands of western Canada and the United States. Sulfur cinquefoil reproduces solely by seed and it is a prolific seed producer. Our 2-yr study (2006, 2007) investigated the optimal timing and intensity of defoliation to decrease aboveground productivity and seed production of sulfur cinquefoil plants on foothill rangeland in southwestern Montana. Each year, 150 sulfur cinquefoil plants within a 430-m2 enclosure were tagged for identification and randomly assigned to one of 15 clipping treatments with 10 plants per treatment. Clipping treatments were conducted at three timings: (preflower [early June], flowering [late June], and seedset [mid-July]) and all possible combinations of timings for a total of seven timing treatments clipped to two stubble heights (7.5 cm or 15 cm), comprising 14 unique treatments. The final (15th) treatment consisted of an unclipped control. Response variables collected at senescence (late July) included aboveground biomass; number of buds, flowers and fruits on each plant; and number and viability of seeds produced. Results indicated that defoliation of sulfur cinquefoil can effectively reduce its yield and seed production. All clipping treatments reduced aboveground biomass of sulfur cinquefoil compared with control plants (P ≤ 0.05), except clipping to 15 cm during preflowering in the wetter year of 2006. Clipping to either 7.5 cm or 15 cm at all times or combinations of timings reduced the number of buds, flowers, fruits, and seeds produced by sulfur cinquefoil (P ≤ 0.05). Viable seed production was reduced 99 to 100% when plants were clipped once to either 7.5 or 15 cm during flowering or seedset. Results suggest that targeted livestock grazing or mowing applied one time per season during flowering or seedset could effectively suppress the biomass production and viable seed production of sulfur cinquefoil.

Most farmers now rely on herbicides and, to a lesser extent, cultivation to control weeds in Glycine max in the Midwest. However, the general public is concerned that widely used herbicides will contaminate surface and groundwater. Alternative ways to control weeds in field crops are needed to reduce or prevent herbicide contamination of surface and groundwater. A new between-row-mowing weed management system that consists of band-applied herbicides over crop rows + two or more between-row mowings was tested in G. max over 6 yr in Missouri. Mowing weeds close to the soil surface two or more times between crop rows killed or suppressed annual grass and broadleaf weeds, chiefly Setaria faberi, Ambrosia artemisiifolia, and Amaranthus spp., if properly timed. Shading by crop canopy closure contributed to weed suppression in this weed management system. G. max yield also could not be distinguished from weed-free check plots and was greater than the weedy check plots. Herbicide use was reduced 50% by banding because only 50% of the field area was sprayed. The between-row-mowing weed management system may have use in environmentally sensitive areas to help reduce soil erosion or water contamination by herbicides.

Defoliation can affect plant competition in at least two ways: negative effects on the performance of defoliated plants and positive effects on nondefoliated plants due to neighbor defoliation. It has been suggested that these effects can alter competitive relations between plants, and this study was designed to test this in an experiment using Crofton weed as the invasive category, and its two neighbors Japanese brome and Dichrocephala as the native category. The plants were grown from seed and potted either singly or in pairs. The pairs consisted either of two conspecific plants (same pairs) or of one plant of each category (mixed pairs). Randomly preselected plants were defoliated twice in the growing season, at 9 wk and again at 5 wk before final biomass sampling. Competition reduced growth by 51 to 78%, with the plants of the native category consistently more affected than the invasive category. When grown singly, Crofton weed grew larger (average 67%) than the native species. As compared with plants grown singly, Crofton weed plants grown in mixed pairs were 40% smaller. The corresponding data for the native species were 78% smaller for Japanese brome and 73% smaller for Dichrocephala. The effects of neighbor defoliation differed among categories and competition. All the nondefoliated plants grown singly were larger than plants grown with a defoliated conspecific neighbor. For the corresponding relationship in mixed pairs, the native plants grown singly were significantly larger than the same native species grown together with a defoliated, invasive neighbor. Defoliation of Crofton weed reduced the growth of native neighbors and increased negative competitive effects on native plants. For the invasive species, however, Crofton weed plants grown in mixed pairs with neighbor defoliation compensated fully for the competitive effects, i.e., they were not significantly different from nondefoliated plants grown singly. Consequently, the effects of defoliation on competition between invasive Crofton weed and its native neighbors may depend on the ability to undertake compensatory growth, and probably on the allelopathic exudation of Crofton weed to herbivory. Our data suggest that herbivory by biocontrols can make an already superior competitor even stronger, especially if the biocontrol agent does not effectively damage or kill the target plant. Owing to complex interactions between competition and defoliation, the indirect effects of herbivory may be more complicated than currently conceived, and understanding the indirect effects of biocontrol agents before their release is crucial.

Alternative methods are needed to control weeds in no-till corn and soybean which minimize herbicide contamination of surface or ground water. The objective of this research was to determine whether between-row (BR) mowing + band-applied herbicide could help reduce herbicide use, without sacrificing summer annual weed control or yield, in no-till soybean and field corn. Glyphosate was applied shortly before or at planting to control emerged winter annual weeds in all treatments. In the BR mowing weed management system, the band-applied soil residual herbicides imazaquin + alachlor in soybean or atrazine + alachlor in corn were applied shortly before or after planting followed by two or more between-row mowings to control summer annual weeds. Annual weeds were first mowed when they were about 8 cm tall and again just before crop canopy closure. Between-row mowing weeds very close to the soil surface two or three times killed or suppressed summer annual grass and broadleaf weeds, chiefly giant foxtail, common cocklebur, and horseweed, when timed properly. The BR mowing weed management system increased yield above a weedy check in these no-till crops. It also controlled weeds and yielded as well as or better than broadcast-applied herbicide at the same rates. Use of soil residual herbicides to control summer annual weeds was reduced 50% by banding because only 50% of the field area was sprayed.

The exotic Brazilian peppertree is a serious invader of both disturbed and natural areas in central and south Florida, forming fast-growing, impenetrable thickets that dominate entire ecosystems. Brazilian peppertree has been targeted for biocontrol, and two defoliating insect species may eventually be released. This study was done to consider the possible effectiveness of defoliating biocontrol agents. The research investigated the effects of different frequencies of defoliation on height, crown diameter, and berry production of young Brazilian peppertrees. All the foliage was manually clipped from 36 trees in field plots once or twice per year for ≥ 1 yr. The effect on berry production of clipping 100% of the leaves from scattered individual branches of one large Brazilian peppertree was ALSo examined. Trees that were completely defoliated five times at 6-mo intervals were significantly smaller and produced significantly fewer fruits than undamaged controls. Plants defoliated one time only, two times in 1 yr, and two times in each of 2 yr were comparable to the undamaged controls. From this simulated herbivory study, we infer that multiple defoliations by insect defoliators have the potential to significantly suppress the growth and fruit production of Brazilian peppertree in Florida.

Introduction. The aim of ourstudy was to analyze the effect of various levels of light availabilityin the fruit microenvironment (induced by summer pruning) on somefruit quality traits. Materials and methods. Two levelsof summer pruning were set: removal of either 30% or 60% of thecanopy leaf area. These treatments were compared with the control(unpruned vines). Two canopy layers (upper and lower) were identifiedin each treatment and in the control vines. At harvest, yield percane and per shoot was recorded in the two canopy layers; fruitfresh weight, flesh firmness, total soluble solid content and totaltitratable acidity were measured. Fruit Total Antioxidant Capacity (TAC)and Total Polyphenols (TPH) were determined at kiwifruit harvestand post-harvest (60 days after storage at 1 °C) in relation totheir position within the canopy (layer) and to the pruning treatmentapplied. Results. Removing 60% of the canopy leaf areainduced higher titratable acidity and flesh firmness of kiwifruitsat harvest; it reduced fruit weight and crop yield, but increasedTAC and TPH compared with kiwifruits of control. Removing 30% leaf areadid not reduce mean fruit weight and crop yield; it increased TACand TPH at harvest by 20%. Post-harvest TAC and TPH evolution didnot differ among treatments. Discussion and conclusion.Pergola-trained kiwifruit vines showed good performances in termsof crop yield and fruit quality when summer pruning reduced the LeafArea Index (LAI) to 3. More intense pruning resulted in a significantincrease in fruit TAC, but reduced fruit weight and crop yield,and delayed softening of pulp.

Responses of plants to herbivory are dependent on the type of damage and the ontogenetic stage of the plant. We compared the effects of stem pruning and defoliation on seedlings of Colophospermum mopane, an ecologically important tree species widely distributed in southern Africa. The growth of 160 greenhouse-grown juveniles were measured for 6-mo after germination and then 6-mo after treatments including 50% defoliation, 100% defoliation, 50% stem pruning and controls. Pruning resulted in 30% reductions in total leaf area, height and biomass. Partial defoliation resulted in 30% reductions in total leaf area and plant biomass. However, complete defoliation resulted in a 30% increase in biomass production, a doubling in leaf and lateral branch number, a 45% reduction in leaf size, and no change in total leaf area. Thus, completely defoliated seedlings showed greater performance than those that were only partially defoliated, indicating that C. mopane has become adapted to the chronic and severe defoliation inflicted by Imbrasia belina caterpillars. Comparison of our results with other studies indicates that C. mopane seedlings are less herbivory-tolerant than adults and that pruning has more negative effects than defoliation. Thus, seedling browsers may constrain recruitment in C. mopane, influencing its population dynamics and abundance.

Damage to Hawaiian Acacia koa forest by Hurricane Iniki was assessed by comparison with our previous measures of stand structure and leaf area index (LAI) at sites along a precipitation/elevation gradient on western Kauai. Reductions in LAI ranged from 29 to 80% and were correlated with pre-hurricane LAI and canopy height. The canopy damage resulted in a large pulse of litter, ranging from 4 to 19 t ha-1 across our study sites. In the first year post-hurricane, LAI recovery and relative increase in stand basal area were negatively correlated with the fraction of canopy LAI lost. The two major overstorey species, A. koa and Metrosiderous polymorpha, were damaged less than the sub-canopy species, Dodonaea viscosa and Psidium guajava. Although D. viscosa and P. guajava were both severely damaged in the hurricane, the alien species P. guajava had much higher survival than the native D. viscosa, both as adults and as seedlings. However, seedling density of P. guajava was limited by low recruitment. At these sites, there was no drastic change in species composition following hurricane disturbance, and forest structure and productivity had recovered to a great degree within 2 y.

Hurricane Gilbert caused disturbance to Jamaican montane rainforests in 1988. This study provides a detailed characterization of landscape-level changes in light below the canopy of these forests after the hurricane. Hemispherical photographs were taken below the forest canopy at four sites at permanent points 1 m above the ground between 7 and 33 mo after the hurricane. For each photograph photosynthetically active radiation (PAR) was computed. PAR declined exponentially in all sites during the period of measurement. During the first 24 mo after the hurricane, PAR beneath the canopy was significantly greater in sites that had been defoliated during the hurricane than in sites where few trees had been defoliated. By 28 mo after the hurricane there was no significant difference in PAR beneath the canopy among the four sites. By 33 mo after the hurricane canopy recovery was nearly complete and PAR was only slightly higher than measurements made before the hurricane. Our results were compared with studies of changes in light environment resulting from treefall gaps and under deciduous canopies. PAR during the first 18 mo after the hurricane was similar to that recorded in small canopy gaps in other forests. Widespread defoliation caused by hurricanes can thus increase PAR beneath the canopy over large areas and consequently opportunities may arise for widespread recruitment of tree species in response to increased light levels.

Fire is a common feature of tropical savannas and it has an important role in the ecology and evolution of the flora. Many woody species in tropical savannas are well adapted to fire and have a vigorous resprouting ability that enables them to survive recurring fire (Frost & Robertson 1987, Hoffmann & Solbrig 2003, Meyer et al. 2005). Studies on the interaction between fire and woody plants in Mediterranean-climate regions have considerably increased our understanding of the subject but relationships between plant attributes (stem diameter and height) and post-fire resprouting ability of southern African savanna woody species are poorly understood. Resprouting is a widespread mechanism by which many plants regenerate after partial or total defoliation (Bellingham & Sparrow 2000, Bond & Midgley 2001).

Brassica rapa grows as a wild and weedy species throughout the world and is the most likely recipient of transgenes from GM oilseed rape. For transgene introgression to occur, the critical step which must be realized, is the formation of an F1 hybrid. Concerns exist that hybrid populations could be more vigorous and competitive compared to the parental species. This study examines the effect of simulated herbivory and interspecific competition on the vegetative and reproductive performance of non-transgenic F1 hybrids and their parental lines. Several vegetative and reproductive performance measures were used to determine the effect of simulated herbivory and competition on the Brassica lines, including leaf length and biomass for herbivory and seedling height and biomass for competition. For defoliation experiments, B. rapa showed little response in terms of leaf length but B. napus and the F1 hybrid responded negatively. Brassica rapa showed elevated biomass responses, but B. napus and the hybrid demonstrated negative responses to defoliation. Defoliation at the cotyledon stage had a slight effect upon final biomass with the F1 hybrid performing significantly worse than B. napus, although seed counts were not significantly different. For the series of competition experiments, hybrids seemed to be more similar to B. rapa in terms of early seedling growth and reproductive measures. The underperformance of hybrid plants when challenged by herbivory and competition, could potentially decrease survivorship and explain the rarity of hybrids in field surveys. However, should transgene introgression occur, the dynamics of hybrids could change radically thus increasing the risk of gene flow from a transgenic oilseed rape crop to the wild recipient.

Abstract This review covers research linking foraging habitat quality for birds to livestock management in lowland farmland. Based on this research we propose a framework for predicting the value of grazing systems to birds. This predictive framework is needed to guide the development of agri-environment measures to address farmland bird declines in pastoral areas. We show that the exacting requirements of declining granivorous birds pose the greatest challenges, while the needs of soil invertebrate feeding species are more easily met.

A severe outbreak of Lepidoptera followed the 1997–98 El Niño Southern Oscillation event, during which the climate in central Panama was unusually dry. The outbreak involved the larvae of at least 12 species of Lepidoptera and occurred at a seasonally dry, deciduous forest site, where extensive background data were available regarding climate, tree species and non-outbreak herbivory levels. Most Lepidoptera were associated with only one or two larval host plant species belonging to the same family, and the majority were monophagous during this study. During the outbreak, caterpillar densities for the major outbreak species averaged 1.6 larvae per young leaf and 0.18 larvae per leaf for leaves of all ages. For canopy trees and lianas, the mean level of leaf damage was 13.8%, ranging from 1–100%. Seven out of 20 tree species sustained most of the damage, with 21–37% of the leaf area consumed. Relative to non-outbreak years, damage levels increased by more than 250% during the outbreak. Single-species outbreaks were observed in other areas with a similar drought, but wetter forests in central Panama did not experience outbreaks during this period. Historically, fewer outbreaks have been reported from tropical forests than from temperate forests, however, similar El Niño-related outbreaks have occurred recently in several other locations throughout the tropics.