Weed seedling emergence is partially dependent on biotic and abiotic conditions directly surrounding the seed. When environmental conditions are appropriate, seed germination and emergence occurs. We studied the impact of seeding depth (surface, 1 to 2, 3 to 4, and 6 to 7 cm) and fluctuating soil moisture regimes (field capacity [FC]–1/3FC–FC; FC–1/6FC–FC) on percent weed emergence in a greenhouse. At FC, wild mustard and field pennycress had the greatest percent emergence when seeds were placed on or near the soil surface, whereas percent emergence of barnyardgrass and round-leaved mallow was unaffected by seeding depth. All the perennials tested had the greatest percent emergence at FC when seeds were placed near or on the soil surface, except for common milkweed which only emerged below the soil surface. When soil moisture levels fluctuated, surface seeds of barnyardgrass, catchweed bedstraw, green foxtail, wheat, and wild oat had less emergence than seeds below the soil surface; field pennycress had increased emergence when the seeds were placed on the surface; and round-leaved mallow and wild mustard emergence was unaffected by seeding depth. The emergence of curly dock, dandelion, and perennial sowthistle was unaffected by seeding depth, whereas foxtail barley and quackgrass emergence was reduced when seeds were placed on the surface and soil moisture fluctuated.

Differences in the depth of weed seedling recruitment due to agronomic management practices, such as reduced tillage, have implications for weed competitive ability and management strategies. Depth of seedling recruitment of Avena fatua, Triticum aestivum, Setaria viridis, Polygonum convolvulus, and Echinochloa crus-galli was measured in situ in 1997 and 1998 prior to seeding (preseeding) and before in-crop spraying (prespray) in a total of 44 zero-tillage and 44 conventional-tillage fields located across approximately 3 million ha of southern Manitoba, Canada. For the monocot species, depth of recruitment was measured from the soil surface to the intact seed coats, which marked the point of germination. For P. convolvulus, a dicot, greenhouse studies were conducted prior to sampling in the field to identify a reliable morphological marker indicating the point of germination. For all species, mean recruitment depth was found to be significantly shallower in zero- vs. conventional-tillage fields and significantly shallower in the preseeding vs. the prespray period. There were relatively few differences in mean recruitment depth among weed species. Within a sampling period and tillage system, for example, the greatest difference in mean recruitment depth between species was less than 1.2 cm, and the maximum mean recruitment depth across species, sampling times, and tillage practice was very shallow (less than 4.2 cm). Locating weed seedling recruitment depth is the first step in characterizing weed seedling recruitment microsites. Results indicate this information should be specific to tillage and sampling time.