A complete coral succession through the Ordovician–Silurian transition in South China reveals an adaptive phase during the Hirnantian glaciation, followed by an early survival phase and finally a late survival phase that persisted into the early Silurian. We demonstrate that a coral assemblage of latest Hirnantian to earliest Silurian age, remarkably similar to those from the Edgewood fauna known from Laurentia, occurs stratigraphically above the typical Hirnantian fauna. This, in combination with other evidence (e.g. brachiopods, lithology and chemostratigraphy), suggests the Edgewood fauna probably post-dated the early–middle Hirnantian glaciation, rather than being coeval with the older glacial-related Hirnantia fauna. Evidence from South China shows that the Edgewood fauna appeared in the very latest Hirnantian and extended into the middle Rhuddanian, considerably younger than previously believed. Such a new correlation necessitates a reassessment of the influence of the end-Ordovician glaciation on biotas. We argue that this major glaciation probably would have substantially affected the ecosystem even in tropical regions, as shown by the development there of the Hirnantia fauna or, alternatively, the presence of a conspicuous stratigraphic hiatus. This suggests a surprisingly rapid biotic recovery during the subsequent postglacial transgression, represented by the flourishing of comparatively diverse shelly faunas (e.g. the Edgewood fauna and the Cathaysiorthis brachiopod fauna) in nearshore shallow water environments from Laurentia to eastern peri-Gondwana terranes or blocks (e.g. South China).

The Hirnantian stratigraphy of Wales is critically assessed against the global evidence for two major Hirnantian synglacial eustatic lowstands. The evidence for two separate lowstands of relative sea level is locally suggestive in shelf sequences but generally only permissive in the basin. Nowhere can the possible candidate lowstands be separated biostratigraphically and thus basinwide correlation that might rule out local tectonic control is impossible to prove. Consistent with palaeolatitude evidence and a situation far removed from the Gondwana margin, there is no facies evidence (e.g. dropstones) for local glacial conditions and the diamictites are of mass-flow origin. Sequence stratigraphic reasoning proves helpful in the correlation of the basinal succession in Central Wales with the shelf sequences along the Tywi Lineament and the Variscan Front, and suggests that the first appearance of Normalograptus persculptus in Wales does not correspond with the base of the persculptus Biozone. Extensive new palaeocurrent data and sedimentological restudy is combined with review of earlier knowledge to argue for significant revisions to previous palaeogeographic maps for ‘generalized lowstand conditions’, notably along the Bala Lineament, the Variscan Front and the Central Wales Inliers.

A pioneer study of the previously unknown δ13C chemostratigraphy in the Ordovician/Silurian boundary interval in eastern Iowa and northeastern Illinois resulted in the discovery of the Hirnantian Isotope Carbon Excursion (HICE). The presence of this major isotope excursion in the Mosalem Formation in Iowa and the Wilhelmi Formation in Illinois, which indicates that the excursion interval in these units is of Hirnantian (latest Ordovician) rather than Early Silurian age, necessitates a revised chronostratigraphic classification of these units. Although the precise level of the Ordovician/Silurian boundary remains somewhat uncertain in the absence of the diagnostic graptolites, it is herein placed in the upper part, but well below the top, of the Mosalem Formation and at the top of the Wilhelmi Formation. During a major regression following the deposition of the Maquoketa Shale, the upper part of the latter clastic unit was in some places deeply eroded, resulting in a topographically dissected landscape with upland areas separated by wide incised valleys. During a subsequent late Hirnantian transgression, these palaeovalleys were gradually filled with marine sediments, but the upland areas were not transgressed until earliest Silurian times. The new chemostratigraphical evidence is in good agreement with the available biostratigraphical data, especially from corals, conodonts, and brachiopods. A preliminary chemostratigraphical study of the presumably coeval Edgewood Group successions in Pike County, northeastern Missouri failed to document any heavy δ13C values characteristic of the HICE and some, or all, of the Hirnantian values obtained there may be diagenetically overprinted.

Understanding the recessional behaviour of ancient pre-Cenozoic ice sheets based on seismic reflection studies is generally difficult through scarcity of data. In North Africa, however, hydrocarbon exploration has produced high quality seismic reflection datasets that permit an analysis of the morphology and internal sedimentary architecture of incisions of Late Ordovician age related to the Hirnantian glaciation. Analysis of a high-resolution 3-D seismic dataset covering a small area in western Libya (the N Murzuq Basin) reveals a sharply defined, WNW–ESE-oriented palaeo-escarpment, with a higher (cliff-forming) western margin and a lower (basin-forming) eastern margin. The palaeo-escarpment defines the western flank of a sub-basin extending up to 60 km in width, known as the Awbari Trough. The escarpment and the trough are interpreted as the morphological expression of a major unconformity dividing pre-glacial sediments below from Late Ordovician (?Hirnantian) glacially related sediments above. Two hypotheses are considered for the origins of both the escarpment and the Awbari Trough: (1) as a tectonic feature such as a half graben that was active during sedimentation and (2) a glacially related palaeotopography, with the latter interpretation preferred, owing to the lack of evidence for syn-sedimentary fault activity. The width of the Awbari Trough compares to the large-scale cross-shelf troughs in modern high latitude settings, such as the Barents Shelf, produced by ice streams. The Awbari Trough was progressively filled in by gravity flow deposits throughout the course of the glaciation, until the initial incision became filled in with sediments during an overall glacial retreat phase and ceased to influence sedimentation patterns. Glacial re-advance across the basin produced a second unconformity observed in seismic data. Above this unconformity, meltwater processes incised a shallow (~ 20 m) and wide (~ 5 km) subglacial tunnel valley. Stabilization of the ice front prior to its ultimate retreat resulted in the deposition of a delta complex prior to the Early Silurian transgression.

The most prominent of the two major global δ13C excursions in the Ordovician, the Hirnantian δ13C excursion (HICE), which is previously recorded from the uppermost Ordovician in a few sections in Nevada, Quebec, Arctic Canada, Baltoscandia, Scotland and China, is documented for the first time from the North American Midcontinent. Samples through the Girardeau Limestone and Leemon Formation in Missouri and Illinois show elevated δ13C values of +4‰ to +5‰. Although not determined precisely, the beginning of the HICE is likely to be in the upper part of the Orchard Creek Shale, and it ends in the upper Leemon Formation. Being extraordinarily useful chronostratigraphically, the presence of the HICE makes it possible to provide a firm dating of the study interval, whose age has long been controversial. Comparison between the study sections and coeval HICE sequences in North America and Europe show striking similarities, especially in sea-level history, indicating that major local lowstands reflect eustatic sea-level changes. A comparison with Hirnantian diamictite successions in North and South Africa and Argentina suggests that these lowstands correspond to two major Gondwanan glacial episodes.

The development of an atypical Hirnantia Fauna in the late Ordovician of Gondwana was coeval with a slow eustatic fall induced by the abstraction of water into a growing ice sheet. This event is dated as early Hirnantian in age and occurred in tandem with the start of a major mass extinction. A tectonic episode in the Caradoc-Ashgill of North Africa differentiated the continental shelf into highs and lows and may have formed the land required for the accumulation of a permanent snow cover. Depositional lows were filled by regressive shallow-marine deposits in the early Hirnantian. During the mid-Hirnantian, advance and retreat of an ice sheet on the continental shelf resulted in the deposition of glaciomarine sediments above these regressive deposits. The demise of an atypical Hirnantia Fauna is attributed to deglaciation and the associated flooding of the continental shelf by a stratified anoxic water column. This glacioeustatic sea-level rise occurred in the late Hirnantian.

Faunas of disarticulated ossicles, particularly columnals, are a potentially important source of data for helping to elucidate the patterns of evolution and extinction shown by pelmatozoans in the late Ordovician. The columnal fauna of the Hirnantian High Mains Formation, Craighead Inlier, Strathclyde, includes the following morphotaxa: Cyclocyclicus (col.) foraminosus sp. nov.; Cyclocyclicus (col.) periogmos sp. nov.; Hypsinomocrinus (col.) lewisi gen. et sp. nov.; Tetragonotetragonalis (col.) girvanensis sp. nov.; and Pentagonopentagonalis (col.) sp. This is the first Hirnantian columnal fauna to be described from the North American province; it is distinctly different from Hirnantian columnal biota described from Wales and Kazakhstan.