A new packrat midden chronology from Playas Valley, southwestern New Mexico, is the first installment of an ongoing effort to reconstruct paleovegetation and paleoclimate in the U.S.A.–Mexico Borderlands. Playas Valley and neighboring basins supported pluvial lakes during full and/or late glacial times. Plant macrofossil and pollen assemblages from nine middens in the Playas Valley allow comparisons of two time intervals: 16,000–10,000 and 4000–0 14C yr B.P. Vegetation along pluvial lake margins consisted of open pinyon–juniper communities dominated by Pinus edulis, Juniperus scopulorum, Juniperus cf. coahuilensis, and a rich understory of C4 annuals and grasses. This summer-flowering understory is also characteristic of modern desert grassland in the Borderlands and indicates at least moderate summer precipitation. P. edulis and J. scopulorum disappeared or were rare in the midden record by 10,670 14C yr B.P. The late Holocene is marked by the arrival of Chihuahuan desert scrub elements and few departures as the vegetation gradually became modern in character. Larrea tridentata appears as late as 2190 14C yr B.P. based on macrofossils, but may have been present as early as 4095 14C yr B.P. based on pollen. Fouquieria splendens, one of the dominant desert species present at the site today, makes its first appearance only in the last millennium. The midden pollen assemblages are difficult to interpret; they lack modern analogs in surface pollen assemblages from stock tanks at different elevations in the Borderlands.

A packrat midden located in the Sierra San Francisco, Baja California Sur, Mexico, dating to ca. 10,200 14C yr B.P., contains remains of California juniper (Juniperus californica) and other taxa now associated with southern California chaparral. California juniper does not occur in the Sierra San Francisco today, although “relict” populations of a few chaparral taxa still occur at higher elevations. This midden record documents the early Holocene occurrence of Baja California coniferous woodland and chaparral vegetation far south of its present distribution or its previously known extent from other fossil records. Based on modern climatic tolerances of California juniper and other taxa, central Baja California experienced a mild Mediterranean-type climate at least 5°–6°C cooler than the climate of today, with at least twice the winter precipitation the region now receives.

Sixty packrat middens were collected in Canyonlands and Grand Canyon National Parks, and these series include sites north of areas that produced previous detailed series from the Colorado Plateau. The exceptionally long time series obtained from each of three sites (> 48,000 14C yr BP to present) include some of the oldest middens yet discovered. Most middens contain a typical late-Wisconsinan glaciation mixture of mesic and xeric taxa, evidence that plant species responded to climate change by range adjustments of elevational distribution based on individual criteria. Differences in elevational range from today for trees and shrubs ranged from no apparent change to as much as 1200 m difference. The oldest middens from Canyonlands NP, however, differ in containing strictly xeric assemblages, including middens incorporating needles of Arizona single-leaf pinyon, far north of its current distribution. Similar-aged middens from the eastern end of Grand Canyon NP contain plants more typical of glacial climates, but also contain fossils of one-seed juniper near its current northern limit in Arizona. Holocene middens reveal the development of modern vegetation assemblages on the Colorado Plateau, recording departures of mesic taxa from low elevation sites, and the arrival of modern dominant components much later.