Palynological results from Julietta Lake currently provide the most direct evidence to support the existence of a glacial refugium for Pinus pumila in mountains of southwestern Beringia. Both percentages and accumulation rates indicate the evergreen shrub survived until at least ∼ 19,000 14C yr BP in the Upper Kolyma region. Percentage data suggest numbers dwindled into the late glaciation, whereas pollen accumulation rates point towards a more rapid demise shortly after ∼ 19,000 14C yr BP. Pinus pumila did not re-establish in any great numbers until ∼ 8100 14C yr BP, despite the local presence ∼ 9800 14C yr BP of Larixdahurica, which shares similar summer temperature requirements. The postglacial thermal maximum (in Beringia ∼ 11,000-9000 14C yr BP) provided Pinus pumila shrubs with equally harsh albeit different conditions for survival than those present during the LGM. Regional records indicate that in this time of maximum warmth Pinus pumila likely sheltered in a second, lower-elevation refugium. Paleoclimatic models and modern ecology suggest that shifts in the nature of seasonal transitions and not only seasonal extremes have played important roles in the history of Pinus pumila over the last ∼ 21,000 14C yr BP.

A sediment core from Smorodinovoye Lake (SML), northeastern Siberia (area to the east of the Verkhoyansk Range) spanning the last 24,000 14C yr indicates that vegetational and climatic changes in the upper Indigirka basin resemble those in eastern Siberia (Lena basin and westward). For example, maximum postglacial summer temperatures at SML probably occurred 6000–4000 14C yr B.P., an age more in accordance with eastern than northeastern records. Larix arrived near the lake by 9600 14C yr B.P., approximately when forests expanded in the east but ca. 1500 14C yr later than forests were established in the neighboring upper Kolyma basin. Paleobotanical data further suggest that Larix possibly migrated southward from populations in the arctic lowlands of eastern Siberia and did not originate from interior refugia of the upper Kolyma basin. Although a Younger Dryas cooling has been noted in eastern Siberia, SML provides the first evidence from the northeast for a similar climatic reversal. Climatic variations seemingly have persisted between the Indigirka and Kolyma basins over at least the last 11,000 14C yr, despite the proximity of the two drainages and the occurrence of major changes in boundary conditions (e.g., seasonal insolation, sea levels) that have influenced other regional climatic patterns.