Sabellidites cambriensis is a tubular non-mineralized metazoan that appears as compressed ribbon-shaped imprints with transverse wrinkling, thick walls and an even tube diameter of up to 3 mm. The distribution of Sabellidites is investigated in three Ediacaran–Cambrian sections on the Digermulen Peninsula in Arctic Norway, spanning the Manndrapselva Member of the Stáhpogieddi Formation and the lower member of the Breidvika Formation. Here, the Ediacaran–Cambrian boundary is located in the lower part of the upper parasequence (third cycle) of the Manndrapselva Member. Specimens of Sabellidites are rare but consistently present close to the lowest level of Treptichnus pedum and upsection, whereas the taxon is common and abundant in the lower part of the lower member of the Breidvika Formation, with an upper record at c. 55 m above the base. The range is comparable with that of the GSSP section in Newfoundland, Canada, establishing Sabellidites as an index fossil for the lowermost Cambrian. In the Manndrapselva Member, Sabellidites co-occurs with the acritarch Granomarginata, indicative of the lowermost Cambrian Granomarginata Zone, whereas in the Breidvika Formation it co-occurs with Asteridium. Sabellidites is widely distributed in Baltica, through the Rovnian and Lontovan regional stages but confined to the Fortunian global stage. In its lower range, Sabellidites is associated with a Treptichnus pedum trace fossil association and a depauperate leiosphaerid acritarch assemblage, followed by a Granomarginata assemblage. In its upper range, Sabellidites co-occurs with acritarchs of the Asteridium–Comasphaeridium Zone and the tubular foraminiferan Platysolenites. In Baltica, Sabellidites is a useful index fossil.

Here we present a detailed accounting of organic microfossils from late Ediacaran sediments of Finland, from the island of Hailuoto (northwest Finnish coast), and the Saarijärvi meteorite impact structure (~170 km northeast of Hailuoto, mainland Finland). Fossils were recovered from fine-grained thermally immature mudstones and siltstones and are preserved in exquisite detail. The majority of recovered forms are sourced from filamentous prokaryotic and protistan-grade organisms forming interwoven microbial mats. Flattened Nostoc-ball-like masses of bundled Siphonophycus filaments are abundant, alongside Rugosoopsis and Palaeolyngbya of probable cyanobacterial origin. Acritarchs include Chuaria, Leiosphaeridia, Symplassosphaeridium and Synsphaeridium. Significantly, rare spine-shaped sclerites of bilaterian origin were recovered, providing new evidence for a nascent bilaterian fauna in the terminal Ediacaran. These findings offer a direct body-fossil insight into Ediacaran mat-forming microbial communities, and demonstrate that alongside trace fossils, detection of a bilaterian fauna prior to the Cambrian might also be sought among the emerging record of small carbonaceous fossils (SCFs).

Detrital zircon U–Pb ages from samples of the Neoproterozoic Visingsö Group, Sweden, yield a maximum depositional age of ≤ 886±9 Ma (2σ). A minimum depositional age is established biochronologically using organic-walled and vase-shaped microfossils present in the upper formation of the Visingsö Group; the upper formation correlates with the Kwagunt Formation of the 780–740 Ma Chuar Group in Arizona, USA, and the lower Mount Harper Group, Yukon, Canada, that is older than 740 Ma. Mineralized scale microfossils of the type recorded from the upper Fifteenmile Group, Yukon, Canada, where they occur in a narrow stratigraphic range and are younger than 788 Ma, are recognized for the first time outside Laurentia. The mineralized scale microfossils in the upper formation of the Visingsö Group seem to have a wider stratigraphic range, and are older than c. 740 Ma. The inferred age range of mineralized scale microfossils is 788–740 Ma. This time interval coincides with the vase-shaped microfossil range because both microfossil groups co-occur. The combined isotopic and biochronologic ages constrain the Visingsö Group to between ≤ 886 and 740 Ma, thus Tonian in age. This is the first robust age determination for the Visingsö Group, which preserves a rich microfossil assemblage of worldwide distribution. The organic and mineralized microorganisms preserved in the Visingsö Group and coeval successions elsewhere document global evolutionary events of auto- and heterotrophic protist radiations that are crucial to the reconstruction of eukaryotic phylogeny based on the fossil record and are useful for the Neoproterozoic chronostratigraphic subdivision.

Until now hemispherical astylospongiid sponges were invariably referred to as Caryospongia juglans var. basiplana Rauff. Renewed investigations have now shown that part of the material should be assigned to a new genus and species, Tympanospongia vankempeni, which is characterised by a system of very irregular canals. These flat-based sponges originate from the Baltic region and occur in two assemblages of silicified Late Ordovician sponges known exclusively as erratics from The Netherlands and northern Germany. These fossils were transported by the River Eridanos, a former drainage system from the Baltic region that filled the North European Basin during the Miocene to Early Pleistocene. Specimens of Tympanospongia vankempeni gen. et sp. nov. also occur in the Upper Pleistocene of Gotland, Sweden. The new sponge described herein principally differs from other genera of the Astylospongiidae found frequently in the erratic sponge assemblages by its irregular system of apochetes which ramify and anastomose commonly.

Compound orchocladine sponges are unusual in the Early Palaeozoic. In Europe, silicified material of Late Ordovician age has hitherto been referred to as Aulocopium aurantium Oswald, 1847 and the invalid Aulocopium compositium Conwentz, 1905. An examination of new material has resulted in the recognition of a new genus, Hydraspongia, with two new species, H. polycephala and H. erecta, and a third new species, Perissocoelia megahabra, to which most specimens can now be assigned. These taxa form part of rich erratic sponge assemblages, which originate from unknown source areas in the Baltic, and have been collected in northern and western Europe from fluvial sandy deposits of the Eridanos River system, which drained the Baltic area from the Middle Miocene to Early Pleistocene.

Altered volcanic ash interbeds (bentonites) in the upper Katian of Baltoscandia indicate significant volcanic activity in neighbouring tectonically active areas. Katian bentonites in the East Baltic can be reliably correlated using sanidine phenocryst composition. Ratios of immobile trace elements TiO2, Nb, Zr and Th to Al2O3 enable extension of the correlations to Scandinavia, where late diagenetic alterations could have caused recrystallization of sanidine phenocrysts. At least seven volcanic eruptions were recognized in Baltoscandian sections. Several bentonites found in deep-sea sediments are absent in shallow-sea sediments, indicating extensive breaks in sedimentation and erosion during late Katian and Hirnantian times. The areal distribution pattern of Katian bentonites in Baltoscandia indicates a volcanic source from the north or northwest (present-day orientation) from the margins of the Iapetus Palaeo-Ocean. Signatures of ultra-high-pressure metamorphism in the Seve Nappe (Central Sweden) and intrusions in the Helgeland Nappe Complex in Central Norway have been proposed as potential sources of the magmas that generated the volcanic ashes deposited in the East Baltic in Katian times. Geochemical similarities between Baltoscandian and Dob's Linn bentonites from southern Scotland suggest a common volcanic source in Katian times.

The Neoproterozoic Hedmark Basin in the Caledonides of South Norway was formed at the western margin of the continent Baltica by rifting 750–600 Ma ago. The margin was destroyed in the Caledonian Orogeny and sedimentary basins translated eastwards. This study uses provenance analysis to map the crustal architecture of the pre-Caledonian SW Baltican margin. Conglomerate clasts and sandstones were sampled from submarine fan, alluvial fan and terrestrial glacigenic sedimentary rocks. Samples were analysed for U–Pb isotopes and clast samples additionally for Lu–Hf isotopes. The clasts are mainly granites c. 960 Ma and 1680 Ma old, coeval with the Sveconorwegian Orogeny and formation of the Palaeoproterozoic Transscandinavian Igneous Belt (TIB). Mesoproterozoic (Sveconorwegian) ages are abundant in the western part of the basin, whereas Palaeoproterozoic ages are common in the east. Lu–Hf isotopes support crustally contaminated source for all clasts linking them to Fennoscandia. Detrital zircon ages of the sandstones can be matched with those from the granitic clasts except for ages within the range 1200–1500 Ma. These ages are typically found in the present-day Telemark, SW Norway. The sandstones and conglomerate clasts in the western part of the Hedmark Basin were sourced from the Sveconorwegian domain in the present SW Norway or its continuation to the present-day NW. The conglomerate clasts in the eastern part of the Hedmark Basin were sourced mainly from the TIB domain or its northwesterly continuation. The Hedmark Basin was initiated within the boundary of two domains in the basement: the TIB and the Sveconorwegian domains.

In the carbonate-siliciclastic strata of West Gondwana (e.g. in the Montagne Noire, France), the aftermath of the mid Languedocian (mid Cambrian) regression is characterized by a late Languedocian major turnover of trilobite families and a Furongian–early Tremadocian radiation related to the stepwise immigration of trilobite invaders from East Gondwana under persistent transgressive conditions. The scarcity of upper Languedocian fossil accumulations in clayey substrates has inspired the sampling of the palaeogeographically most distal parts of the Iberian Chains (Spain), where diagenetic dissolution of ubiquitous hexactinellid sponge spicules has favoured the formation of siliceous concretions. These have yielded the trilobites Peronopsis cf. insignis, Oidalagnostus trispinifer, Proampyx difformis (= Proampyx aculeatus), Bailiaspis? glabrata (= Holocephalina agrauloides, by ontogeny), Holasaphus cf. centropyge and a paradoxidid gen. et sp. indet. Despite preservation and sampling biases, the identification of this taphonomic window in offshore clayey substrates of West Gondwana allows the recognition of a strong biogeographical link with Baltica, and the correlation of the global Guzhangian Stage and the Solenopleura? brachymetopa Zone of Scandinavia with part of the Mediterranean upper Languedocian Substage.

The trilobite fauna of the upper Ordovician (middle Katian) Pyle Mountain Argillite comprises a mixture of abundant mesopelagic cyclopygids and other pelagic taxa and a benthic fauna dominated by trilobites lacking eyes. Such faunas were widespread in deep water environments around Gondwana and terranes derived from that continent throughout Ordovician time but this is the only known record of such a fauna from North America and thus from Laurentia. It probably reflects a major sea level rise (the ‘Linearis drowning events’) as does the development of coeval cyclopygid-dominated deep water trilobite faunas in terranes that were marginal to Laurentia and are now preserved in Ireland and Scotland. The Pyle Mountain Argillite trilobite fauna occurs with a deep water Foliomena brachiopod fauna and comprises 22 species. Pelagic trilobites (mostly cyclopygids) constitute 36% of the preserved sclerites, and 45% of the fauna is the remains of trilobites lacking eyes, including one new species, Dindymene whittingtoni sp. nov. Three species of cyclopygid are present, belonging in Cyclopyge, Symphysops and Microparia (Heterocyclopyge). Cyclopygids are widely thought to have been stratified in the water column in life and thus their taxonomic diversity reflects the relative depths of the sea-beds on which their remains accumulated. A tabulation of middle and upper Katian cyclopygid-bearing faunas from several palaeoplates and terranes arranged on the basis of increasing numbers of cyclopygid genera allows an assessment of the relative depth ranges of the associated benthic taxa. The Pyle Mountain Argillite fauna lies towards the deeper end of this depth spectrum.

The lower–middle Cambrian transitional interval of Scania is largely represented by condensed limestone beds, lithostratigraphically grouped in the Gislöv Formation (1–5.7 m thick), and the Forsemölla and Exsulans Limestone beds (lower part of the Alum Shale Formation, up to 4 m thick). The strata display a combination of skeletal carbonate productivity, episodic nucleation of phosphate hardground nodules, and polyphase reworking recorded on a platform bordering the NW corner of Baltica. The shell accumulations can be subdivided into three deepening-upward parasequences, separated by distinct erosive unconformities. The parasequences correspond biostratigraphically to the Holmia kjerulfi, Ornamentaspis? linnarssoni and Ptychagnostus gibbus zones, the latter two generally being separated by a stratigraphic gap that includes the middle Cambrian Acadoparadoxides oelandicus Superzone. Except for the Exsulans Limestone, the carbonates reflect development of a prolific epibenthic biota, dominated by filter-feeding nonreefal chancelloriid–echinoderm–sponge meadows, rich in trilobites and brachiopods, and which were subjected to high-energy conditions. The absence of microbial mats or veneers encrusting the erosive surfaces of these event-concentration low-relief shoal complexes may be related to long hiatal episodes resulting in microboring proliferation. High levels of nutrient supply resulted in high primary productivity, eutrophic conditions, glauconite precipitation, phosphogenesis (in some case microbially mediated) and microendolithic infestation. An early-diagenetic mildly reducing environment is suggested by the presence of authigenic (subsequently reworked) pyrite, which contrasts with the syndepositional normal oxygenated conditions reflected by macroburrowing and the abundance of benthic fossils.

The Severnaya Zemlya Archipelago, located close to the continental edge of the Kara Shelf in the Russian high Arctic, represents, together with northern Tajmyr, the exposed Neoproterozoic and Palaeozoic part of the North Kara Terrane. This terrane has been interpreted as an independent microcontinent or part of a larger entity, such as Arctida or Baltica, prior to collision with Siberia in Late Carboniferous time. A major stratigraphic break, the Kan'on (canyon) River Unconformity, separates folded Late Cambrian from Early Ordovician successions in one area, October Revolution Island. New geochronological U–Th–Pb ion-microprobe data on volcanic and intrusive rocks from this island constrain the age of an important magmatic episode in the earliest Ordovician. A tuff, in association with Tremadocian fossils, overlying the Kan'on River Unconformity, has been dated to 489.5 ± 2.7 Ma. The youngest rocks beneath the unconformity are of the Peltura minor Zone, and the latter has been dated previously, in western Avalonia, to 490.1+1.7−0.9 Ma. Thus, little time is available for the tectonic episode recorded by the unconformity, and the similarities in radiometric dates may indicate problems with the correlation of faunal markers for the Cambrian–Ordovician boundary across palaeo-continents. The other extrusive and intrusive rocks which have been related to Early Ordovician rifting in the Severnaya Zemlya area yield ages from 489 Ma to 475 Ma. An undeformed granite, cutting folded Neoproterozoic successions on neighbouring Bol'shevik Island has been dated to 342 ± 3.6 Ma and 343.5 ± 4.1 Ma (Early Carboniferous), in accord with evidence elsewhere of Carboniferous strata unconformably overlying the folded older successions. This evidence conflicts with the common interpretation that the structure of the Severnaya Zemlya Archipelago originated during the collision of the North Kara Terrane with Siberia in Late Carboniferous time. An alternative interpretation is that Severnaya Zemlya was located in the Baltica foreland of the Caledonide Orogen and that the eastward-migrating deformation of the foreland basin reached the area of the archipelago in latest Devonian to Early Carboniferous time. This affinity of the North Kara Terrane to Baltica is further supported by 540–560 Ma xenocrysts in Ordovician intrusions on October Revolution Island, an age which is characteristic of the Timanide margin of Baltica.

Erosional outliers are an important component of the Ludlow (Late Silurian) stratigraphy of Gotland, Sweden. However, due to the poor degree of exposure, outcrop studies have not revealed a detailed section from any of these outliers. The first complete stratigraphy of such an erosional remnant, the Burgen outlier, is presented here, based on the recently recovered Burgen-1 drillcore. Stratigraphical units encountered are, from oldest to youngest, the uppermost Hemse Group (>35 m), Eke Formation (0.4 m), Burgsvik Formation (3.4 m) and the basal Hamra Formation (12 m preserved). Based on carbon stable isotopes and lithological correlation to nearby outcrops that are zoned by conodonts, it can be concluded that the most profound late Ludlow sedimentary changes took place shortly after the onset of the globally recognized Lau Event (δ13C values reach 8.7 ‰ in the lower Hamra Formation). Sedimentary changes include development of discontinuity surfaces, widespread occurrence of an intraclastic conglomerate, increased microbial activity (Rothpletzella and Wetheredella), and influx of clastics to the basin. These changes are linked to substantial base-level changes during the event. A general comparison with the Prague basin suggests contemporaneous sedimentary changes on Baltica and on cratonic elements from peri-Gondwana during the event.

Biogenic apatite preserved in 148 samples of conodonts and organophosphatic-shelled brachiopods from Cambrian through Ordovician successions of the Baltoscandian Basin (Baltica Plate) preserves a sensitive record of early Palaeozoic sea-water chemistry interpreted via neodymium isotope ratios. Consistent ${\uvarepsilon}$Nd(t) values of −9.6 to −8.3 for Lower to Middle Cambrian samples suggest no significant lateral or temporal variation across the region. Average Upper Cambrian values are −7.2 to −7.7. Sedimentary analysis suggests that the influence of continental weathering from Baltica as a major source of radiogenic Nd was negligible. Ordovician samples show a rise to −5 to −6 in the early Arenig, early–mid Llanvirn and late Caradoc. Sea-water mixing from the southeast Iapetus Ocean was a constant factor throughout Cambrian–Ordovician times. The rise reflects erosion of obducted volcanic arc complexes along the Caledonian margin, and probably also relates to pollution of the Baltica sector of Iapetus from the approaching Avalonia Plate. Patterns of evolutionary biodiversity and palaeobiogeographical linkages support the geochemical signatures in interpreting the tectonic history of the region. Extinction of lingulate brachiopod faunas in the Tremadoc, followed by subsequent recovery and emergence of benthic assemblages typical of the Ordovician Evolutionary Fauna in the Billingen–early Volkhov regional stages coincide with significant changes in geochemical characteristics of water masses across the Baltoscandian basin. The early and mid Ordovician (Arenig to Llandeilo) brachiopod faunas of the North Estonian Confacies Belt are characterized by high endemism and low turnover rates, whereas increased immigration resulted in the extinction of a number of local lineages in the late Llanvirn. From the mid Caradoc to mid Ashgill, when Baltica was drifting on a course to collide eventually with Avalonia and gradually approach Laurentia, brachiopod assemblages were characterized by higher turnover rates. At the same time they gradually became more cosmopolitan and less influenced by the invasion of new faunas.

Conodont faunas from the Eoplacognathus pseudoplanus Zone in five main areas of Middle Ordovician rocks in Sweden are described and discussed and compared with faunas of comparable age from neighbouring areas. The investigation is based on 118 samples which together yielded 175 114 conodont elements. The Swedish conodont faunas are intermediate between shallower-water faunas from the eastern part of Baltica and deeper-water faunas towards the west. Correlation with the Yangtze platform of south-central China is fairly easy, and the occurrence of key taxa, such as Histiodella, makes it also possible to correlate some levels with Laurentia. Although faunal lists for the Swedish localities are almost identical, the conodont taxa have their relative abundance maxima in separate areas: species of Protopanderodus and Periodon are most common in deeper-water settings, Eoplacognathus, Baltoniodus and Microzarkodina characterize intermediate areas, and Semiacontiodus is typical of shallow subtidal environments

The available data from Newfoundland, the British Isles and Scandinavia suggest that by late Ordovician–early Silurian times the ocean separating Laurentia from Eastern Avalonia and Baltica had partly closed with the consumption of intervening oceanic crust. Marine seaways, however, persisted until the middle or late Silurian. Phases of crustal transtension and transpression, predominantly under a major sinistral shear couple, occurred throughout the Silurian and early Devonian until the remnant Iapetus Ocean was completely destroyed. The most appropriate Recent plate tectonic models for Silurian sedimentation between Eastern Avalonia and Laurentia are probably the deep-marine foreland basins between Timor and the northern Australian margin, or between Taiwan and mainland China.